Tachytes

Tachytes distinctus, copyright xpda.

Belongs within: Larrini.

Tachytes: crickets face death from above
Published 21 April 2016
Tachytes species feeding, copyright Stephen Cresswell.

Most people who are not entomologists assume that ‘ants’, ‘bees’ and ‘wasps’ are all mutually exclusive groups of animals. But while bees and ants are distinct from each other, they are both really distinctive subgroups of wasps. It is not difficult to find guides on the interwebs purporting to tell you the differences between a bee and a ‘wasp’, but many of the points usually cited will not apply to all wasps (usually what is intended is the differences between a bee and a social wasp, which is the type of non-bee, non-ant wasp most likely to make itself known to humans). Some wasps can look very similar to bees indeed.

Diagram of Tachytes ocelli, from here.

Tachytes is a large, cosmopolitan genus of not-quite-bees, including more than 250 species found on all continents except Antarctica. Its members are robust and hairy, and one would have to look very closely to spot the features marking it as a non-bee (such as the point that its hairs, though numerous, are not branched in the manner of a true bee). Tachytes is classified in the Crabronidae, the family of wasps believed to be most closely related to true bees (other crabronids have been featured in earlier posts: here, here and here). Like other crabronids, adult Tachytes are pollinators, feeding on nectar from flowers. Tachytes species can be separated from most other crabronids by the shape of the ocelli. As well as the two large compound eyes with their multiple lenses on either side of the head, many insects have three small single-lens eyes, called the ocelli, on the top of the head (some insects have fewer or no ocelli). When all three ocelli are present, they are arranged in a triangle with one at the front and two at the rear. In Tachytes, the rear ocelli are present but deformed (I presume non-functional, though I don’t actually know). They are reduced to a pair of scar-lines shaped roughly like a comma, or the upper part of a question mark without the dot.

Female Tachytes sinensis sinensis with katydid prey, from here.

Like many other wasps, female Tachytes dig burrows in which they they sequester other insects in a paralysed state to provide food for their young (the big change between bees and other wasps was the provision of their larvae with a plant-derived food source such as honey or pollen instead of animal matter). The size of the burrow varies from species to species, but some are quite extensive: the North American species T. praedator digs a burrow with a centimetre-wide entrance about a metre long, with a 70 cm down-shaft followed by a horizontal run of about a foot (Lin 1967). This species digs at night, thus presumably both avoiding the heat of the day and reducing the risk of detection by predators or parasites. The top of the burrow is usually marked by a heap comprised of the removed soil from its digging; in some species, the female will close over the top of the burrow when not actively digging. A series of cells are constructed branching from the end section of the burrow; each of these cells is filled with enough prey to feed one larva, then an egg is laid in the cell and its entrance sealed. Most Tachytes species feed their larvae with Orthoptera such as crickets, grasshoppers or katydids; the exact type preferred differs between species, though I get the impression of a general correlation between the size of the Tachytes species and the size of its preferred prey. At least two exceptional Tachytes species from central Asia (T. ambidens and T. bidens) are known to supply their larvae with small caterpillars rather than orthopterans. The prey is carried by the female held by her legs, with the antennae clasped by the mandibles. Hunting behaviour has been described for two species from North America, T. intermedius and T. mergus, that stock their nest with pygmy mole crickets (Tridactylidae). Females of these species walk along the ground tapping at it with their antennae. When one locates a cricket in its underground burrow, she quickly digs downwards in an attempt to grab the cricket by its head with her mandibles and haul it out. Even if the cricket hears her coming and attempts to flee from its burrow, it may not escape. The wasp moves so quickly that she is often able to grab the cricket in mid-air as it leaps for freedom. These two Tachytes species also differ from others in the genus in that the crickets are sequestered in the nest not fully paralysed: it has been noted that mole crickets recovered from their nests are still quite mobile and even able to jump.

Systematics of Tachytes
Tachytes Panzer 1806 (see below for synonymy)BM76
|--T. (Tachytes)B78
| |--T. obsoletus groupB78
| | |--T. (T.) obsoletus (Rossi 1792)GOY03 [=Apis obsoletaBM76, *Tachyplena obsoletaBM76, *Tachyptera obsoletaBM76]
| | | |--T. o. obsoletusBM76
| | | |--T. o. occidentalis Pulawski 1962BM76
| | | `--T. o. tricoloratus (Turton 1802)BM76 (see below for synonymy)
| | |--T. (T.) flagellarius Bohart 1978B78
| | |--T. (T.) stangei Bohart 1978B78
| | `--T. (T.) varians (Fabricius 1804)B78, BM76 [=Liris variansBM76]
| `--T. leprieurii groupB78
| |--T. (T.) hades Schrottky 1903B78, BM76 [incl. T. minos Shrottky 1903BM76]
| |--T. (T.) jucundus Smith 1856 [incl. T. fulvipes Schrottky 1903 non Smith 1856]B78
| |--T. (T.) leprieurii (Spinola 1841)B78, BM76 [=Lyrops leprieuriiBM76; incl. T. apiformis Smith 1856BM76]
| |--T. (T.) palatus Bohart 1978B78
| |--T. (T.) rufalaris Bohart 1978B78
| |--T. (T.) rugulosus Bohart 1978B78
| `--T. (T.) schlingeri Bohart 1978B78
`--T. (Tachynana Banks 1942)B87, BM76
| i. s.: T. (T.) abdominalis (Say 1823)L67, BM76 [=Larra abdominalisBM76]
| T. (T.) coloratus Bohart 1978B78
| T. (T.) concinnus Smith 1856 [incl. T. argyrofacies Strand 1910]B78
| T. (T.) menkei Bohart 1978B78
| T. (T.) micantipygus Strand 1910B78, BM76
| T. (T.) rhodogaster Bohart 1978B78
| T. (T.) richardsi Bohart 1978B78
| T. (T.) willinki Bohart 1978B78
| T. (T.) zuliae Bohart 1978B78
|--T. chrysopyga groupB78
| |--T. (T.) chrysopyga (Spinola 1841) (see below for synonymy)B78
| | |--T. c. chrysopygaBM76
| | |--T. c. argentipes Smith 1856 [incl. T. insularis Cresson 1865]BM76
| | |--T. c. obscurus Cresson 1872 (see below for synonymy)BM76
| | `--T. c. staegeri Dahlbom 1845BM76
| |--T. (T.) catarinae Bohart 1978B78
| |--T. (T.) excellens Cameron 1912B78, BM76
| |--T. (T.) pubescens Bohart 1978B78
| |--T. (T.) roraimae Bohart 1978B78
| `--T. (T.) simulans Smith 1873B78, BM76
|--T. amazonus groupB78
| |--T. (T.) amazonus Smith 1856B78, BM76 (see below for synonymy)
| |--T. (T.) fritzi Bohart 1978B78
| |--T. (T.) rubioi Bohart 1978B78
| |--T. (T.) ustulatus Bohart 1978B78
| `--T. (T.) venezuelae Bohart 1978B78
|--T. (T.) ambidens Kohl 1884B78, BM76 [incl. T. caucasicus Radoszkowski 1886BM76]
|--T. chilensis groupB78
| |--T. (T.) andesae Bohart 1978B78
| |--T. (T.) chilensis (Spinola 1851) (see below for synonymy)B78
| `--T. (T.) imperialis Saussure 1867B78, BM76 [incl. T. saussurei Reed 1894BM76]
|--T. fraternus groupB78
| |--T. (T.) alvarengai Bohart 1978B78
| |--T. (T.) aurovestitus Bohart 1978B78
| |--T. (T.) catamarcae Bohart 1978B78
| |--T. (T.) chrysocercus Rohwer 1911 [incl. T. apache Banks 1942]BM76
| |--T. (T.) costalis Taschenberg 1870B78, BM76
| |--T. (T.) distinctus Smith 1856B78, BM76 (see below for synonymy)
| | |--T. d. distinctusBM76
| | `--T. d. bimini Krombein 1953BM76
| |--T. (T.) fraternoides Bohart 1978B78
| |--T. (T.) fraternus Taschenberg 1870 (see below for synonymy)B78
| |--T. (T.) frontalis Smith 1873B78
| `--T. (T.) vardyi Bohart 1978B78
|--T. (T.) integer Gussakovskij 1932B78, BM76
|--T. minutior groupB78
| |--T. (T.) lissinus Bohart 1978B78
| |--T. (T.) minutior Bohart 1978B78
| `--T. (T.) politus Bohart 1978B78
|--T. nitidiusculus groupB78
| |--T. (T.) nitidiusculus (Smith 1856)B78, BM76 [=Larrada nitidiusculaBM76, Tachysphex nitidiusculusBM76]
| |--T. (T.) absidatus Bohart 1978B78
| |--T. (T.) ornatipes Cameron 1889B78, BM76
| |--T. (T.) quinquedens Bohart 1978B78
| |--T. (T.) rhododactylus Taschenberg 1870B78, BM76 [incl. T. callosus Kohl 1892BM76]
| |--T. (T.) sexdens Bohart 1978B78
| `--T. (T.) villegasi Bohart 1978B78
|--T. (T.) pygmaeus Kohl 1888B78, BM76 (see below for synonymy)
`--T. setosus groupB78
|--T. (T.) pretiosus Cameron 1912B78, BM76
|--T. (T.) setiger Kohl 1898 [=T. setigera]B78
`--T. (T.) setosus Taschenberg 1870B78, BM76

Tachytes incertae sedis:
T. abercornensis Arnold 1959BM76
T. admirabilis Turner 1916BM76
T. aeneus Saunders 1910BM76
T. aestuans Turner 1916BM76
T. agadiriensis Nadig 1933BM76
T. alacris Arnold 1960BM76
T. albonotatus Walker 1871BM76
T. alfierii Pulawski 1962BM76
T. andreniformis Cameron 1889 [=T. anthreniformis Dalla Torre 1897]BM76
T. angustiverticisLCL08
T. antillarum Cameron 1906BM76
T. apiformis Smith 1856BM76
T. approximatus Turner 1908BM76
T. archaeophilus Pulawski 1962BM76
T. argenteovestitus Cameron 1910BM76
T. argenteus Gussakovskij 1932 [incl. T. radoszkowskii Beaumont 1936]BM76
T. argyreus (Smith 1856) (see below for synonymy)BM76
T. argyropis Saussure 1887BM76
T. asiagenes Pulawski 1962BM76
T. assamensis Cameron 1904BM76
T. associatus Turner 1917BM76
T. astatiformis Tsuneki 1963BM76
T. astutus Nurse 1909 [incl. T. shirozui Tsuneki 1966]BM76
T. aureocinctus Cameron 1905BM76
T. aurichalceus Kohl 1891BM76
T. auricomans Bradley 1919BM76
T. aurifex Smith 1858BM76
T. auripes Berland 1942BM76
‘Lyrops’ auritaR26
T. aurulentus (Fabricius 1804) (see below for synonymy)BM76
T. badius Banks 1942BM76
T. bakeri Williams 1928BM76
T. basilicus (Guérin-Méneville 1844) (see below for synonymy)BM76
T. beludzhistanicus Gussakovskij 1932BM76
T. bidens Gussakovskij 1952BM76
T. bimetallicus Turner 1917BM76
T. birkmanni Rohwer 1909 [incl. T. atomus Banks 1942, T. obscuranus Rohwer 1909]BM76
T. biskrensis Saunders 1910BM76
T. borneanus Cameron 1902 [incl. T. banoensis Rohwer 1919]BM76
T. braunsi Turner 1917BM76
T. bredoi Arnold 1947BM76
T. brevipennis Cameron 1900 [incl. T. maculitarsis Cameron 1900]BM76
T. brevis Walker 1871BM76
T. brunneomarginatus Arnold 1947BM76
T. brunneus Pulawski 1962BM76
T. bulawayoensis Bischoff 1913BM76
T. californicus Bohart 1962BM76
T. cameronianus Morice 1897BM76
T. carinatus Berland 1945BM76
T. cataractae Arnold 1923BM76
T. celsissimus Turner 1917BM76
T. cephalotes Walker 1871BM76
T. ceratophorus Pulawski 1962BM76
T. chelatus Bohart 1962BM76
T. chivensis Pulawski 1962BM76
T. chudeaui Berland 1942BM76
|--T. c. chudeauiBM76
`--T. c. niger Berland 1942 non Sphex niger Fabricius 1775 (not preoc. if in dif. gen.)BM76
T. cinerascens Arnold 1935BM76
T. codonocarpi Pulawski 1975BM76
T. columbiae Fox 1892BM76
T. comberi Turner 1917 (see below for synonymy)BM76
T. compactus Arnold 1951BM76
T. confusus Arnold 1923BM76
T. contractus Walker 1871BM76
T. copiosus Arnold 1945BM76
T. corniger Gussakovskij 1952 [=T. cornigera; incl. T. trichopus Pulawski 1962]BM76
T. crassus Patton 1881BM76
T. danae Arnold 1923BM76
T. decoratus Walker 1871BM76
T. decorsei Berland 1942BM76
T. dichrous Smith 1856 [=*Holotachytes dichrous; incl. Larrada ferox Smith 1873]BM76
|--T. d. dichrousBM76
`--T. d. hospes Bingham 1898BM76
T. dilaticornis Turner 1916BM76
T. diodontus Pulawski 1962BM76
T. discrepans Arnold 1951BM76
T. dispersus Turner 1916BM76
T. disputabilis Turner 1917BM76
T. distanti Turner 1917BM76
T. diversicornis Turner 1918 [incl. T. griseolus Arnold 1951, T. rufitibialis Arnold 1951]BM76
T. dubiosus Tsuneki 1974BM76
T. ermineus Banks 1942BM76
T. erynnis Turner 1917BM76
T. etruscus (Rossi 1790)GOY03 (see below for synonymy)
|--T. e. etruscusLCL08
`--T. e. sibiricus Gussakovskij 1932LCL08, BM76
T. europaeus Kohl 1884BM76
T. eurous Pualwski 1962BM76
T. exclusus Turner 1917BM76
T. exornatus Fox 1893BM76
T. falciger Arnold 1951BM76
T. famelicus Pulawski 1962BM76
T. fatalis Turner 1916BM76
T. fervens Smith 1873BM76
T. fervidus Smith 1856BM76
T. fidelis Pulawski 1962BM76
T. flagellatus Nurse 1903BM76
T. flavocinereus Arnold 1945BM76
T. floridanus Rohwer 1920 [incl. T. comanche Banks 1942, T. foxi Banks 1942]BM76
T. formosissimus Turner 1908BM76
T. freygessneri Kohl 1802LCL08
T. fontalis Smith 1873BM76
T. fruticis Tsuneki 1964BM76
|--T. f. fruticisBM76
`--T. f. taianus Tsuneki 1974BM76
T. fucatus Arnold 1951 [incl. T. serapis Pulawski 1962]BM76
T. fulviventris Cresson 1865 [incl. T. caelebs Patton 1879, T. coelebs Dalla Torre 1897]BM76
|--T. f. fulviventrisBM76
`--T. f. rossi Bohart 1962BM76
T. fulvopilosus Cameron 1904 (see below for synonymy)BM76
T. gondarabai (Guiglia 1943) [=Tachysphex gondarabai]BM76
T. gracilicornis Arnold 1944BM76
T. guatemalensis Cameron 1889 [incl. T. coxalis Patton 1892]BM76
T. habilis Turner 1917BM76
T. hainanensis Wu & Li 2006LCL08
T. hamiltoni Turner 1917 [incl. T. dubitatus Rohwer 1909, T. harpax Patton 1881]BM76
T. hengchunensis Tsuneki 1967BM76
T. hirsutus Smith 1856BM76
T. illabefactus Pulawski 1962BM76
T. indicus Dalla Torre 1897 [incl. T. ornatipes Cameron 1889 non Cameron 1889]BM76
T. indifferens Arnold 1945BM76
T. indulgens Cheesman 1937BM76
T. inexorabilis Turner 1917BM76
T. instabilis Turner 1917BM76
T. intermedius (Viereck 1906)BM76 (see below for synonymy)
T. interstitialis Cameron 1900BM76
T. irreverens Cheesman 1937BM76
T. irritabilis Turner 1917BM76
T. kristenseni Turner 1917BM76
T. labilis Turner 1917BM76
T. lachesis Turner 1917BM76
T. lamborni Arnold 1934BM76
T. lamentabilis Arnold 1951BM76
T. lanuginosus Pulawski 1962BM76
T. latiforns Tsuneki 1964BM76
T. lepidus Arnold 1929BM76
T. levantinus Pulawski 1962BM76
T. lingnaui Arnold 1933BM76
T. longirostris Arnold 1947BM76
T. lugubris Walker 1871BM76
T. maculicornis Saunders 1910BM76
T. maerens Turner 1917 [=T. moerens Arnold 1923]BM76
T. magellanicus Williams 1928BM76
T. maindroni Berland 1942BM76
T. manjikuli Tsuneki 1963BM76
T. maroccanus Pulawski 1962 [incl. T. aegyptiacus Pulawski 1962]BM76
T. marshalli Turner 1912 [=*Calotachytes marshalli]BM76
T. matronalis Dahlbom 1845GOY03 [incl. T. obesus Kohl 1884BM76]
T. megaerus Turner 1917BM76
T. melancholicus Arnold 1923BM76
T. memnon Turner 1916BM76
T. meraukensis Cameron 1911BM76
T. mergus Fox 1892 [=*Tachyoides mergus; incl. Tachyoides ariellus Banks 1942, Tachytes minor Rohwer 1909]BM76
T. midas Arnold 1929BM76
T. mirus Kohl 1894 [incl. T. pulchrivestitus Cameron 1908]BM76
T. mitis Turner 1916BM76
T. modestus Smith 1856 [incl. T. dilwara Nurse 1903, T. maculipennis Cameron 1904]BM76
|--T. m. modestusBM76
`--T. m. neglectus Turner 1917BM76
T. monetarius Smith 1856BM76
T. mongolicus Tsuneki 1972BM76
T. mutilloides Walker 1871BM76
T. nasicornis Gussakovskij 1952 [incl. T. rhinoceros Gussakovskij 1952]BM76
T. natalensis Saussure 1854BM76
T. nebulosusB78
T. nevadensis Bohart 1962BM76
T. nigrescens Berland 1942BM76
T. nigroannulatus Bischoff 1913 [incl. T. glabriusculus Arnold 1923]BM76
|--T. n. nigroannulatusBM76
`--T. n. lichtenburgensis Arnold 1923BM76
T. nigropilosellus (Cameron 1910) [=Liris nigropilosella; incl. T. gigas Bischoff 1913]BM76
T. niloticus Turner 1918 [incl. T. curiosus Gussakovskij 1952]BM76
T. nipponicus Tsuneki 1964BM76
T. nitidulus (Fabricius 1793) [=Crabro nitidulus]BM76
T. nomarches Pulawski 1962BM76
T. notabilis Turner 1917BM76
T. nudiventris Turner 1917BM76
T. obductus Fox 1892BM76 [=T. (Tachynana) obductusL67]
T. observabilis Kohl 1894BM76
T. oppositus Turner 1917BM76
T. ‘opulentus’ Nurse 1909 non Lepeletier 1845BM76
T. oviventris Saussure 1891BM76
T. pallidiventris Arnold 1947BM76
T. ‘panzeri’ Dufour 1841 non Vander Linden 1829GOY03
|--T. p. panzeriLCL08
`--T. p. orientis Pulawski 1962LCL08, BM76 [=T. europaeus orientisBM76]
T. parvus Fox 1892 [incl. T. arizonicus Banks 1942, T. pattoni Banks 1942]BM76
T. pennsylvanicus Banks 1921BM76
T. pepticus (Say 1837) (see below for synonymy)BM76
T. perornatus Turner 1917BM76
T. picticornis Arnold 1945BM76
T. plagiatus Walker 1871BM76
T. plutocraticus Turner 1910BM76
T. popovi Pulawski 1962BM76
T. praedator Fox 1892BM76 [=T. (Tachyplena) praedatorL67]
T. praestabilis Turner 1917BM76
T. priesneri Pulawski 1962BM76
T. procerus Costa 1882 [incl. T. incognitus Pulawski 1962]BM76
T. pulchricornis Turner 1917 [incl. T. kolaensis Turner 1917]BM76
T. punctuosus Arnold 1923BM76
T. rarus Arnold 1960BM76
T. relucens Turner 1916BM76
T. repandus (Fabricius 1787) [=Crabro repandus, Vespa repanda]BM76
T. rhodesianus Bischoff 1913BM76
T. rostratus Berland 1942BM76
T. rubellus Turner 1908BM76
T. rufipalpis Cameron 1904BM76
T. ‘rufipes’ Berland 1942 nec Aichinger 1870 nec Reed 1894BM76
T. rufiscutis Turner 1918BM76
T. rufomarginatus Arnold 1945BM76
T. sacricola Pulawski 1962BM76
T. sagani Guiglia 1943BM76
T. saharicus Pulawski 1962BM76
T. salvus Kohl 1906BM76
T. saundersii Bingham 1897 (see below for synonymy)BM76
|--T. s. saundersiiBM76
`--T. s. suluensis Williams 1928BM76
T. sayi Banks 1942 [incl. T. brevipilis Banks 1942, T. hesperus Banks 1942]BM76
T. sculleni Bohart 1962BM76
T. sedulus Smith 1860BM76
T. senegalensis Berland 1942BM76
T. separabilis Turner 1917BM76
T. sheppardi Arnold 1940BM76
T. silverlocki Turner 1917BM76
T. silvicola Williams 1928BM76
T. silvicoloides Williams 1928BM76
T. simillimus Schulthess 1926BM76
T. simulatrix Turner 1917BM76
T. sinensis Smith 1856 [incl. T. japonicus Kohl 1888, T. rothneyi Cameron 1889]BM76
|--T. s. sinensisBM76
|--T. s. fundatus Rohwer 1911BM76
`--T. s. yaeyamanus Tsuneki 1972BM76
T. sinuatus Pulawski 1962BM76
T. sjoestedti Cameron 1908BM76
|--T. s. sjoestedtiBM76
|--T. s. formosus Arnold 1923BM76
`--T. s. karrooensis Arnold 1923BM76
T. spatiosus Li, Cai & Li 2008LCL08
T. spatulatus Fox 1892 [incl. T. basirufus Rohwer 1909, T. utahensis Banks 1942]BM76
T. sulcatus Turner 1916BM76
T. surigensis Williams 1928BM76
T. tabrobane Cameron 1900 [incl. T. proximus Nurse 1903]BM76
T. tachyrrhostus Saussure 1854BM76
T. tarsalis (Spinola 1838) (see below for synonymy)BM76
T. tassilicus Pulawski 1962BM76
T. testaceinerva Cameron 1908BM76
T. tomentosus Kohl 1891BM76
T. toyensis Tsuneki 1971BM76
T. transvaalensis Cameron 1905BM76
T. trichopygus Pulawski 1962BM76
T. tricinctus (Fabricius 1804) [=Liris tricincta; incl. T. cubensis Cresson 1865]BM76
T. trigonalis Saussure 1867BM76
T. turneri Arnold 1923BM76
T. ugandensis Turner 1917BM76
T. vagus Radoszkowski 1877BM76
T. validus Cresson 1872 (see below for synonymy)BM76
T. velox Smith 1856 [incl. T. neavei Turner 1917]BM76
T. versatilis Turner 1917BM76
T. vestitus (Smith 1873) (see below for synonymy)BM76
T. vicinus Cameron 1889BM76
T. vischnu Cameron 1889 [=T. virchu, T. vischu Dalla Torre 1897, T. vishnu Bingham 1897]BM76
T. volubilis Turner 1917BM76
T. xenoferus Rohwer 1911BM76
T. yerburyi Bingham 1897BM76
T. yunnanensis Li, Cai & Li 2008LCL08

Tachytes Panzer 1806 [incl. Calotachytes Turner 1917, Holotachytes Turner 1917, Lyrops Illiger 1807, Tachyoides Banks 1942, Tachyplena Banks 1942, Tachyptera Dahlbom 1843 non Berge 1842]BM76

Tachytes argyreus (Smith 1856) [=Larrada argyrea; incl. Tachysphex debilis Pérez 1907, Tachytes melanopygus Costa 1893]BM76

Tachytes aurulentus (Fabricius 1804) [=Larra aurulenta; incl. T. duplicatus Rohwer 1920, T. mandibularis Patton 1881, T. propinquus Rohwer 1909]BM76

Tachytes basilicus (Guérin-Méneville 1844) [=Lyrops basilicus; incl. T. guichardi Arnold 1951, T. superbiens Morice 1911]BM76

Tachytes comberi Turner 1917 [incl. T. atlanticus Berland 1942, T. patrizii Guiglia 1932, T. tricinctus Pérez 1907 non Liris tricincta Fabricius 1804]BM76

Tachytes etruscus (Rossi 1790)GOY03 [=Andrena etruscaBM76, Larra etruscaL49, *Lyrops etruscusBM76; incl. Tachytes argentatus Brullé 1833BM76]

Tachytes fulvopilosus Cameron 1904 [incl. T. andreniformis Cameron 1902 non Cameron 1889, T. fulvovestitus Cameron 1904]BM76

Tachytes intermedius (Viereck 1906)BM76 [=Tachysphex intermediusBM76; incl. Tachytes amiculus Banks 1942BM76, Tachytes austerus Mickel 1916BM76, Tachytes maestus Mickel 1916BM76, Tachytes minutus Rohwer 1909BM76, T. (Tachynana) minutusL67]

Tachytes pepticus (Say 1837) [=Lyrops pepticus; incl. T. cressoni Banks 1942, T. inferioris Banks 1942, T. sericatus Cresson 1872]BM76

Tachytes saundersii Bingham 1897 [incl. T. formosanus Tsuneki 1966, T. shiva Nurse 1903, T. varipilosus Cameron 1905]BM76

Tachytes tarsalis (Spinola 1838) [=Lyrops tarsalis; incl. Larrada tarsata Smith 1856 non T. tarsatus Smith 1856, T. turcomanica Radoszkowski 1893]BM76

Tachytes validus Cresson 1872 [incl. T. belfragei Banks 1942, T. breviventris Fox 1892, T. calcaratiformis Rohwer 1909, T. calcaratus Fox 1892, T. coloradensis Banks 1942, T. quadrifasciatus Dreisbach 1948]BM76

Tachytes vestitus (Smith 1873) [=Larrada vestita; incl. Tachysphex tarsalis Dalla Torre 1897 non Lyrops tarsalis Spinola 1838, Tachytes tarsatus Smith 1856 non Larra tarsata Say 1823]BM76

Tachytes (Tachynana) amazonus Smith 1856B78, BM76 [=Tachytes amazonumB78; incl. Tachytes anisitsi Strand 1910BM76, Tachytes clypeatus Taschenberg 1870BM76, Tachytes dives Holmberg 1884 non Lepeletier 1845BM76, Tachytes fervens Smith 1873BM76, Tachytes fiebrigi Brèthes 1909BM76, Tachytes holmbergii Dalla Torre 1897BM76, Tachytes nigricaudus Brèthes 1909BM76, Tachytes rufoannulatus Strand 1910BM76, Tachytes rufofasciatus Cresson 1872BM76, Tachysphex rufomaculatus Cameron 1889BM76, Tachytes scalaris Taschenberg 1870BM76, Tachytes tuberculatus Giner Marí 1944B78]

Tachytes (Tachynana) chilensis (Spinola 1851) [=Larra chilensis; incl. T. dichrous Herbst 1921 non Smith 1856, L. gayi Spinola 1851, T. mimeticus Schrottky 1911, T. spinolae Reed 1894]B78

Tachytes (Tachynana) chrysopyga (Spinola 1841) [=Lyrops chrysopyga; incl. T. argenticrus Dalla Torre 1897, T. argentipes Cameron 1889 non Smith 1856, T. columbianus Saussure 1867]B78

Tachytes (Tachynana) chrysopyga obscurus Cresson 1872 [=*Tachynana obscura; incl. Tachytes hirsutifrons Banks 1942, Tachytes texanus Cresson 1872]BM76

Tachytes (Tachynana) distinctus Smith 1856B78, BM76 [incl. T. austrinus Banks 1942BM76, T. contractus Fox 1892 non Walker 1871BM76, T. elongatus Cresson 1872BM76, Larrada fulvipes Smith 1856BM76, T. seminole Banks 1942BM76, T. yucatanensis Cameron 1889BM76]

Tachytes (Tachynana) fraternus Taschenberg 1870 [incl. T. ametina Cameron 1912, T. asuncionis Strand 1910, T. ferrugineipes Cameron 1889 non Lepeletier 1845]B78

Tachytes (Tachynana) pygmaeus Kohl 1888B78, BM76 [incl. T. basalis Cameron 1889BM76, T. calvus Turner 1929BM76, T. seminudus Arnold 1951BM76]

Tachytes (Tachytes) obsoletus tricoloratus (Turton 1802)BM76 [=Sphex tricoloratusBM76; incl. T. tingitanus Pate 1937BM76, S. tricolor Fabricius 1793 non Schrank 1781BM76, Larra tricolorL49, Lyrops tricolorR26, Pompilus tricolorBM76, *Tachytes tricolorBM76]

*Type species of generic name indicated

References

[B78] Bohart, R. M. 1978. Tachytes of South America (Hymenoptera, Sphecidae, Larrinae). Transactions of the American Entomological Society 104 (3–4): 435–505.

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[GOY03] Gayubo, S. F., H. Özbek & E. Yıldırım. 2003. A contribution to the knowledge of Spheciformes of Turkey: tribes Astatini, Dinetini, Larrini, Palarini and Trypoxylini (Hymenoptera: Apoidea: Crabronidae). Zoology in the Middle East 29: 83–92.

[LCL08] Li, T., W. Cai & Q. Li. 2008. Two new species and five new records of the genus Tachytes Panzer (Hymenoptera, Crabronidae) from China, with a key to the Chinese species. Deutsche Entomologische Zeitschrift 55 (1): 153–159.

[L67] Lin, C. S. 1967. Nesting behavior of Tachytes (Tachyplena) praedator Fox, with a review of the biology of the genus (Hymenoptera: Sphecidae: Larrinae). American Midland Naturalist 77 (1): 241–245.

[L49] Lucas, H. 1849. Exploration Scientifique de l’Algérie pendant les années 1840, 1841, 1842 publiée par ordre du gouvernement et avec le concours d’une commission académique. Sciences physiques. Zoologie. II. Histoire naturelle des animaux articulés. Troisième partie. Insectes (suite). Imprimerie Nationale: Paris.

[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.

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