Therevinae

Ozodiceromyia sp., possibly O. notata, copyright Yurika Alexander.

Belongs within: Therevidae.

The Therevinae are a widespread group of stiletto flies characterised primarily by features of the genitalia. Members include the diverse North American genus Ozodiceromyia, characterised by areas of reduced pubescence on the frons, thorax and/or abdomen, with the frons usually being shiny black in females and having a small shiny black area in males. Anabarhynchus and related genera have a setose metanepisternum, abundant occipital macrosetae loosely arranged in several rows, and lack setulae on vein R1 (Gaimari & Webb 2009).

Characters (from Winterton et al. 2001): Male genitalia with dorsal apodeme of parameral sheath free from gonocoxal apodeme; ventral apodeme of parameral sheath simple. Female genitalia with acanthophorites well developed; intersegmental membrane between sternite 8 and furca sclerotised; tergite 8 joined to tergite 9+10 by sclerotised bridge; two spermathecae; spermathecal sac present. Pupa with alar spine present. Larva with labial beard present; anterior maxillary palp large.

<==TherevinaeGW09
    `--CycloteliniGW09
         |--Anolinga longiventrisGW09
         |--Breviperna placidaGW09
         |--ColeianaGW09
         |--CrebrasetaGW09
         |--NesonanaGW09
         |--SpiracolisGW09
         |--Cyclotelus Walker 1856B90
         |    |--C. bellusGW09
         |    |--C. nigrifronsGW09
         |    |--C. pictipennisGW09
         |    |--C. rufiventrisGW09
         |    `--C. sumichrastiGW09
         `--Ozodiceromyia Bigot 1889GW09, B90
              |--*O. mexicana Bigot 1889B90
              |--O. argentiferaGW09
              |--O. costalisBT04
              |--O. notataGW09
              `--O. signatipennisGW09
Therevinae incertae sedis:
  Anabarhynchus Macquart 1847WY01, B90
    |--A. calceatusCM91
    |--A. niger Bigot 1889B90
    `--A. tristis Bigot 1889B90
  IrwiniellaWY01
  MegatherevaWY01
  PlatycarenumWY01
  Brachylinga cinereaIL81, GW09
  Thereva Latreille 1796 [incl. Exapata Macquart 1840]B90
    |--*T. plebeiaL02, R26 [=Bibio plebejaL02]
    |--T. anilisG20
    |--T. bipunctataK01
    |--T. fucataGW09
    |--T. fulvaR26
    |--T. marginulaK01
    `--T. nobilitataRD77
  Psilocephala Bigot in Zetterstedt 1842WY01, B90
    |--P. aldrichiiE66
    |--P. electrella Cockerell 1920P92
    |--P. hypogaeaIL81
    |--P. indica Bigot 1889B90
    |--P. macrochaeta Bigot 1889B90
    |--P. pilosula Bigot 1889B90
    |--P. rufipes Bigot 1889B90
    `--P. scudderiIL81

*Type species of generic name indicated

References

[B90] Bigot, J.-M.-F. 1890. Diptères nouveaux ou peu connus. 35e partie. Annales de la Société Entomologique de France, 6e série 9: 313–328.

[BT04] Bybee, S. M., S. D. Taylor, C. R. Nelson & M. F. Whiting. 2004. A phylogeny of robber flies (Diptera: Asilidae) at the subfamilial level: molecular evidence. Molecular Phylogenetics and Evolution 30: 789–797.

[CM91] Colless, D. H., & D. K. McAlpine. 1991. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 717–786. Melbourne University Press: Carlton (Victoria).

[E66] Evans, H. E. 1966. The Comparative Ethology and Evolution of the Sand Wasps. Harvard University Press: Cambridge (Massachusetts).

[GW09] Gaimari, S. D., & D. W. Webb. 2009. Therevidae (stiletto flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 633–647. NRC Research Press: Ottawa.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[IL81] Irwin, M. E., & L. Lyneborg. 1981. Therevidae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 513–523. Research Branch, Agriculture Canada.

[K01] Kertész, K. 1901. Legyek [Dipteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 179–201. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.

[RD77] Richards, O. W., & R. G. Davies. 1977. Imms’ General Textbook of Entomology 10th ed. vol. 2. Classification and Biology. Chapman and Hall: London.

[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.

[WY01] Winterton, S. L., L. Yang, B. M. Wiegmann & D. K. Yeates. 2001. Phylogenetic revision of Agapophytinae subf. n. (Diptera: Therevidae) based on molecular and morphological evidence. Systematic Entomology 26: 173–211.

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