Thynninae

Upa tridentata, from Lopes Justino et al. (2013).

Belongs within: Euaculeata.
Contains: Agriomyia, Zeleboria, Lophocheilus, Aeolothynnus, Thynnus.

The Thynninae are a group of wasps exhibiting extreme sexual dimorphism. Males are winged and have the apex of the hypopygium produced and often spinose; females are wingless and often lack functional ocelli. Males often have the hypopygium produced and spinose or dentate (Kimsey 2004).

<==Thynninae
| i. s.: Oncorhinus xanthospilus [incl. Thynnus gravidus Westw. 1844]T08
| RhytidogasterT08
| |--R. discrepans Turner 1908T08
| `--R. pyxidatus Turner 1908T08
| BraunsomeriaB28
|--ThynniniK04
| |--Acanthothynnus sannaeK04
| |--Arthrothynnus mulleriK04
| |--Belothynnus fuscocostalisK04
| |--Bifidothynnus wubiniensisK04
| |--Campylothynnus flavopictusK04
| |--ChilothynnusK04
| |--DythynnusK04
| |--Doratithynnus doddiK04
| |--Elidothynnus melleusK04
| |--Epactiothynnus abductorK04
| |--Gymnothynnus gilbertiK04
| |--Hathynnus pygmaeusK04
| |--Leiothynnus mackayensisK04
| |--Leptothynnus purpureipennisK04
| |--Lestricothynnus illidgeriK04
| |--Megalothynnus friederichiK04
| |--Neozeleboria [incl. Aulacothynnus]K04
| |--Pentazeleboria agnataK04
| |--PogonothynnusK04
| |--Tachynoides huntianusK04
| |--Zythynnus vespoidesK04
| |--Tmesothynnus [incl. Aspidothynnus]K04
| | `--T. zeleboriK04
| |--Catocheilus Guér. 1842K04, T08 [incl. Hemithynnus Ashm. 1903K04, T08]
| | `--*C. klugii Guérin 1842 (see below for synonymy)T08
| |--Guerinius Ash. 1903K04, T08 [incl. TachynothynnusK04]
| | `--G. picipesK04
| |--Iswaroides Ashm. 1903K04, T08 [incl. AsthenothynnusK04]
| | `--*I. koebelei Ashm. 1903T08
| |--Tachyphron [incl. Takyomyia]K04
| | `--T. evelinaeK04
| |--Zaspilothynnus Ashm. 1903K04, T08
| | `--Z. fenestrusK04
| |--AriphronK04
| | |--A. blandulusK04
| | `--A. rigidulusK04
| |--BeithynnusK04
| | |--B. amplusK04
| | `--B. solarisK04
| |--EncopothynnusK04
| | |--E. atrifaciesK04
| | `--E. neoatrifaciesK04
| |--OncorhinothynnusK04
| | |--O. xanthorrhoeiK04
| | `--O. xanthospilusK04
| |--Psammothynnus Ashm. 1903T08
| | |--*P. depressus (Westw. 1844) (see below for synonymy)T08
| | `--P. fulvopilosus (Sm. 1879) [=Thynnus fulvopilosus; incl. Rhagigaster rugosus Sm. 1879]T08
| |--TachynomiaK04
| | |--T. abdominalisK04
| | `--T. imbellis Turner 1908T08
| |--Thynnoides Guér. 1839T08
| | |--T. gracilis (Westw. 1844)K04, T08 (see below for synonymy)
| | |--T. rufiabdominalis Rayment 1935R35
| | `--T. unifasciatusK04
| |--Caetrathynnus Brown 2000B00
| | |--*C. greggi Brown 2000B00
| | |--C. galbinus Brown 2000B00
| | `--C. wesselensis Brown 2000B00
| |--Nitidothynnus Brown 2000B00
| | |--*N. purdiei Brown 2000B00
| | |--N. ebeneus Brown 2000B00
| | `--N. spinulus Brown 2000B00
| |--Procerothynnus Brown 2000B00
| | |--*P. centralianus Brown 2000B00
| | |--P. arnhemicus Brown 2000B00
| | `--P. carpentarianus Brown 2000B00
| |--Phymatothynnus Turner 1908T08
| | |--*P. monilicornis (Sm. 1859) [=Thynnus (Agriomyia) monilicornis]T08
| | |--P. aratus (Turner 1908) [=Thynnus (Phymatothynnus) aratus]T08
| | |--P. distinctus [=Lophocheilus distinctus, Thynnus (Phymatothynnus) distinctus]T08
| | `--P. nitidus (Sm. 1859) [=Thynnus (Agriomyia) nitidus]T08
| |--Macrothynnus Turner 1908T08
| | |--*M. simillimus (Sm. 1859) [=Thynnus simillimus; incl. T. molitor Sm. 1859]T08
| | |--M. dilatatus (Sm. 1859) [=Thynnus dilatatus]T08
| | |--M. friederici (D.T. 1897) (see below for synonymy)T08
| | |--M. insignis (Sm. 1859) [=Thynnus insignis]T08
| | `--M. poultoni (Turner 1908) [=Thynnus (Macrothynnus) poultoni]T08
| |--AgriomyiaK04
| |--ZeleboriaK04
| |--LophocheilusK04
| |--AeolothynnusK04
| `--ThynnusK04
|--ScotaeniniK04
| |--Anodontyra quadrizonataK04
| |--Glottynnus laraK04
| |--Ornepetes nigricepsK04
| |--Parelaphroptera santacrucianaK04
| |--Pseudelaphroptera tricolorK04
| |--RostrynnusK04
| | |--R. tarsatusK04
| | `--R. tricolorK04
| `--ScotaenaK04
| |--S. fastuosa (Sm. 1879) [=Thynnus fastuosus]T08
| |--S. flavovariegata (Sm. 1879) [=Thynnus flavovariegatus]T08
| |--S. geniseiK04
| |--S. horniK04
| |--S. polistoidesK04
| `--S. trifasciata [incl. Thynnus decorus]T08
|--ElaphropteriniK04
| |--AmblysomaK04
| |--Ammodromus frontalisK04
| |--Argenthynnus impresusK04
| |--Atopothynnus unidensK04
| |--Brethynnus infernusK04
| |--Chrysothynnus incaK04
| |--Dolichothynnus pastorisK04
| |--EucyrtothynnusK04
| |--MerithynnusK04
| |--Mesothynnus gratiosusK04
| |--SpilothynnusK04
| | |--S. exsectusK04
| | `--S. laetusK04
| |--TelephoromyiaK04
| |--UpaK04
| | |--*U. impressiceps [=Scotaena impressiceps]K96
| | |--U. bidentata Kimsey 1996K96
| | |--U. ligulata Kimsey 1996K96
| | |--U. longispina Kimsey 1996K96
| | |--U. nasuta Kimsey 1996K96
| | |--U. porteri Kimsey 1996K96
| | `--U. tridentata Kimsey 1996K96
| |--Zeena aethiopsK04
| `--ElaphropteraK04
| |--E. scoliaeformisK04
| `--E. vulpinaK04
`--RhagigasteriniK04
|--Aelurus nigrofasciatusK04
|--Eirone transversusK04
|--DimorphothynnusK04
| |--D. lecheriK04
| `--D. obtususK04
`--RhagigasterK04
|--R. lyelliK04
|--R. obtususT08
`--R. unicolorZS10

*Catocheilus klugii Guérin 1842 [=Thynnus (*Catocheilus) klugii; incl. T. diversus Sm. 1859, T. perplexus Sm. 1879]T08

Macrothynnus friederici (D.T. 1897) [=Thynnus friederici; incl. T. klugii Westw. 1844 non Catocheilus klugii Guérin 1842]T08

*Psammothynnus depressus (Westw. 1844) [=Thynnus (Agriomyia) depressus, Zeleboria depressa; incl. T. trisulcatus Sm. 1859]T08

Thynnoides gracilis (Westw. 1844)K04, T08 [=Thynnus (Thynnoides) gracilisT08; incl. Thynnus (T.) bidens Sauss. 1867T08, Thynnus dallatorrei Schulz 1906T08, Thynnus viduus Sauss. 1867T08]

*Type species of generic name indicated

References

[B28] Betrem, J. G. 1928. Monographie der Indo-Australischen Scoliiden mit zoogeographischen Betrachtungen. H. Veenman & Zonen: Wageningen.

[B00] Brown, G. R. 2000. Caetrathynnus, Nitidothynnus and Procerothynnus, new genera of Thynninae (Hymenoptera: Tiphiidae) from northern Australia. Beagle, Records of the Museums and Art Galleries of the Northern Territory 16: 107–122.

[K96] Kimsey, L. S. 1996. Revision of the South American thynnine genus Upa (Hymenoptera: Tiphiidae). Proceedings of the Entomological Society of Washington 98 (1): 55–63.

[K04] Kimsey, L. S. 2004. Illustrated keys to genera of the male wasps in the subfamily Thynninae (Hymenoptera: Tiphiidae). Proceedings of the Entomological Society of Washington 106 (3): 571–585.

[R35] Rayment, T. 1935. A Cluster of Bees: Sixty essays on the life-histories of Australian bees, with specific descriptions of over 100 new species. Endeavour Press: Sydney.

[T08] Turner, R. E. 1908. A revision of the Thynnidae of Australia. [Hymenoptera]. Part II. Proceedings of the Linnean Society of New South Wales 33: 70–256.

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