Erechthias acontistes, copyright CSIRO.

Belongs within: Ditrysia.
Contains: Tinea.

The Tineidae are a group of small to medium-sized moths that rest with the body flat and the wings tented, and often tend to run when disturbed rather than fly. They are characterised by a more or less vestigial proboscis and antennae with a single row of scales on each segment. Several species are economic pests on animal fibres and grain such as the common clothes moth Tineola bisselliella (Nielsen & Common 1991).

Within the Tineidae, members of the Tineinae have an apical half-rosette of lateral bristles on the second segment of the labial palp in addition to a row of ventral bristles (Nielsen & Common 1991). Hieroxestinae have smooth scales on the head forming a ‘brow-ridge’, a strongly oblique frons, long and slender scapes, and long spurs (Nielsen & Common 1991).

Characters (from Nielsen & Common 1991): Small to medium sized; head usually rough, with hair-scales, or smooth scaled; chaetosemata and ocelli absent; antennae simple filiform, scape usually with pecten; proboscis usually short, long in some genera; maxillary palps five-segmented and folded, or reduced; labial palps drooping, porrect or ascending, second segment usually with lateral bristles; epiphysis usually present, spurs 0-2-4, hind tibiae never smooth scaled; fore wing with chorda and M-stem often present in cell, M frequently forked, Rs to costa or apex; ovipositor usually telescopic. Eggs oval and slightly flattened. Larva sometimes case-bearing, usually feeding on dried animal or vegetable matter, or on fungi, with 0–6 stemmata on each side of head, crochets in uniserial circle or ellipse. Pupa usually with two transverse rows of spines on terga, rarely with one; segments 4–7 movable in male, 4–6 in female; in larval shelter or cocoon.

|--Harmaclona [Harmacloninae]KP19
|--Nemapogon [Nemapogoninae]NC91
| `--N. granellaNC91
| |--Setomorpha rutellaNC91
| |--Lindera tessellatellaD96, NC91
| `--Prosetomorpha Davis 1996D96
| `--*P. falcata Davis 1996D96
| |--TinissaNC91
| |--Morophaga clonodesNC91
| `--ScardiaC70
| |--S. australasiellaP27b
| |--S. boletiP01
| |--S. clonodesP27b
| `--S. primaevaP27b
|--Erechthias [Erechthiinae]NC91
| |--E. acontistesNC91
| |--E. acrodinaL27
| |--E. charadotaL27
| |--E. externellaM27
| |--E. fulguritellaL27
| |--E. lychnopa Meyrick 1927M27
| |--E. mystacinellaNC91
| `--E. symmachaC70
| |--TimaeaNC91
| |--MimoscopaNC91
| |--GeronthaNC91
| |--Edosa [Perissomasticinae]NC91
| | `--E. meliphanesNC91
| `--MoerarchisNC91
| |--M. australasiellaZS10
| `--M. clathrataNC91
| |--HieroxestisP27b
| | |--H. hapsimachaL27
| | `--H. omoscopaP27b
| `--Opogona [Oinophilidae]NC91
| |--O. comptellaC70
| |--O. glycyphagaNC91
| |--O. micranthesP27b
| |--O. omoscopaNC91
| `--O. punctata (Walsingham 1900) [=Dendroneura punctata]JG19
| |--Tineola bisselliellaNC91, WRM02
| |--AcridotarsaNC91
| |--TineaGE05
| |--Niditinea fuscellaNC91
| `--MonopisNC91
| |--M. argillaceaNC91
| |--M. chrysogrammaNC91
| |--M. ethelellaP27b
| `--M. rusticellaRD77
|--Ankistrophorus giganteus Druce 1901D01
|--Thysanosedes salvini Druce 1901D01
| |--F. echinon Druce 1901D01
| `--F. garleppi Druce 1901D01
|--A. arcei Druce 1901D01
|--A. numidia Druce 1901D01
|--A. perrensi Druce 1901D01
|--A. punctata Druce 1901D01
`--A. whitelyi Druce 1901D01

Tineidae incertae sedis:
Prothinodes grammocosmaP27b
Sagephora phortegellaP27b
Archyala terraneaP27b
Trichophaga tapetzellaRD77
Obesoceras abchasicusYS10
|--L. epixylaP27b
`--L. howesiL27
|--M. araneosaP27a
|--M. fragilis Philpott 1927P27a
`--M. strigulataP27b
Achrea grisellaM86
|--C. acontistesC70
`--C. mystacinellaC70
Monopibaltia ignitellaRJ93
Tineolamina aurellaP92
Palpula pyropella [incl. Tinea bicostella]L49
Helina flammella [=Tinea flammella; incl. T. formosella]L49
|--H. characterellaL49
|--H. cicutellaL49
|--H. dictamnellaL49
`--H. jugurthella Lucas 1849E12

*Type species of generic name indicated


[C70] Common, I. F. B. 1970. Lepidoptera (moths and butterflies). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 765–866. Melbourne University Press.

[C74] Common, I. F. B. 1974. Lepidoptera (moths and butterflies). In: CSIRO. The Insects of Australia: A textbook for students and research workers. Supplement 1974 pp. 98–107. Melbourne University Press.

[D96] Davis, D. R. 1996. Neotropical Tineidae, VI: Prosetomorpha falcata, a new genus and species of Setomorphinae (Lepidoptera: Tineoidea) from Colombia associated with curculionid galleries in stems of Solanum. Proceedings of the Entomological Society of Washington 98 (2): 173–187.

[D01] Druce, H. 1901. Descriptions of some new species of Lepidoptera from east Africa and tropical America. Annals and Magazine of Natural History, series 7, 7: 432–444.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[JG19] James, D. J., P. T. Green, W. F. Humphreys & J. C. Z. Woinarski. 2019. Endemic species of Christmas Island, Indian Ocean. Records of the Western Australian Museum 34 (2): 55–114.

[KP19] Kawahara, A. Y., D. Plotkin, M. Espeland, K. Meusemann, E. F. A. Toussaint, A. Donath, F. Gimnich, P. B. Frandsen, A. Zwick, M. dos Reis, J. R. Barber, R. S. Peters, S. Liu, X. Zhou, C. Mayer, L. Podsiadlowski, C. Storer, J. E. Yack, B. Misof & J. W. Breinholt. 2019. Phylogenomics reveals the evolutionary timing and pattern of butterflies and moths. Proceedings of the National Academy of Sciences of the USA 116 (45): 22657–22663.

[L27] Lindsay, S. 1927. A list of the Lepidoptera of Deans Bush, Riccarton, Canterbury. Transactions and Proceedings of the New Zealand Institute 57: 693–696.

[L49] Lucas, H. 1849. Exploration Scientifique de l’Algérie pendant les années 1840, 1841, 1842 publiée par ordre du gouvernement et avec le concours d’une commission académique. Sciences physiques. Zoologie. II. Histoire naturelle des animaux articulés. Troisième partie. Insectes (suite). Imprimerie Nationale: Paris.

[M86] Masters, G. 1886. Notes and exhibits. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 164.

[M27] Meyrick, E. 1927. Descriptions of New Zealand Lepidoptera. Transactions and Proceedings of the New Zealand Institute 57: 697–702.

[NC91] Nielsen, E. S., & I. F. B. Common. 1991. Lepidoptera (moths and butterflies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 817–915. Melbourne University Press: Carlton (Victoria).

[P01] Pável, J. 1901. Lepkék [Lepidopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 171–177. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[P27a] Philpott, A. 1927a. New Zealand Lepidoptera: notes and descriptions. Transactions and Proceedings of the New Zealand Institute 57: 703–709.

[P27b] Philpott, A. 1927b. The maxillae in the Lepidoptera. Transactions and Proceedings of the New Zealand Institute 57: 721-746.

[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.

[RD77] Richards, O. W., & R. G. Davies. 1977. Imms’ General Textbook of Entomology 10th ed. vol. 2. Classification and Biology. Chapman and Hall: London.

[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.

[WRM02] Wiegmann, B. M., J. C. Regier & C. Mitter. 2002. Combined molecular and morphological evidence on the phylogeny of the earliest lepidopteran lineages. Zoologica Scripta 31 (1): 67–81.

[YS10] Yefremova, Z. A., & I. S. Strakhova. 2010. A review of the species of the genus Elasmus Westwood (Hymenoptera, Eulophidae) from Russia and neighboring countries. Entomologicheskoe Obozrenie 89 (3): 634–661.

[ZS10] Zborowski, P., & R. Storey. 2010. A Field Guide to Insects in Australia 3rd ed. Reed New Holland: Sydney.

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