Titanopterida

Reconstruction of Gigatitan sp., from Gorochov & Rasnitsyn (2002).

Belongs within: Pantcholmanvissiida.

The Titanopterida are very large, raptorial insects known from the Triassic of Asia and Australia. Béthoux (2007) identified titanopteridans as derived from basal Orthoptera, implying a loss of jumping abilities in this lineage. They were defined by Béthoux (2007) by the synapomorphy of forewings with veins CuPaα• (the posterior branch of CuPaα from its first branching) + CuPaβ and CuPb having the same point of origin.

<==Titanopterida [Mesotitanida, Mesotitanina, Mesotitanini, Mesotitanoptera, Titanoptera]
    |  i. s.: Permotitan Gorochov 2004B07
    |           `--P. vladimiri Gorochov 2004B07
    |--Gigatitan Sharov 1968B07 [incl. Nanotitan Sharov 1968B07; GigatitanidaeGR02]
    |    |--G. extensus (Sharov 1968) [=Nanotitan extensus]B07
    |    |--G. magnificus (Sharov 1968)B07
    |    `--G. vulgaris Sharov 1968B07
    |--ParatitanidaeG03
    |    |--Microtitan Gorochov 2003G03
    |    |    `--*M. zherichini Gorochov 2003G03
    |    `--Paratitan Sharov 1968G03
    |         |--P. bispeculum Gorochov 2003G03
    |         |--P. intermedius Gorochov 2003G03
    |         |--P. latispeculum Gorochov 2003G03
    |         |--P. libelluloides Sharov 1968G03
    |         |--P. longispeculum Gorochov 2003G03
    |         |--P. modestus Gorochov 2003G03
    |         |--P. ovalis Sharov 1968G03
    |         |--P. reductus Gorochov 2003G03
    |         `--P. venosus Gorochov 2003G03
    `--MesotitanidaeG03
         |--Prototitan Sharov 1968 [Prototitaninae]G03
         |    |--*P. primitivus Sharov 1968G03
         |    `--P. sharovi Gorochov 2003G03
         `--Mesotitaninae [Clathrotitanidae]G03
              |--Ultratitan Sharov 1968G03
              |    `--U. superior Sharov 1968G03
              |--Mesotitanodes Sharov 1968G03
              |    |--M. similis (Sharov 1968) [=Prototitan similis]G03
              |    `--M. tillyardi Sharov 1968G03
              `--Mesotitan Tillyard 1916B07 [incl. Clathrotitan McKeown 1937G03]
                   |--M. giganteus Tillyard 1916B07
                   |--M. libelluloides (Sharov 1968)B07
                   |--M. ovalis (Sharov 1968)B07
                   |--M. primitivus (Sharov 1968)B07
                   |--M. scullyi Tillyard 1925G03 (see below for synonymy)
                   `--M. superior (Sharov 1968)B07

Mesotitan scullyi Tillyard 1925G03 [=Clatrotitan scullyiF71; incl. Clathrotitan andersoni McKeown 1937G03, M. tillyardi McKeown 1937G03]

*Type species of generic name indicated

References

[B07] Béthoux, O. 2007. Cladotypic taxonomy applied: titanopterans are orthopterans. Arthropod Systematics and Phylogeny 65 (2): 135–156.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.

[G03] Gorochov, A. V. 2003. New and little known Mesotitanidae and Paratitanidae (Titanoptera) from the Triassic of Kyrgyzstan. Paleontologicheskii Zhurnal 2003 (4): 62–69 (translated: Paleontological Journal 37 (4): 400–406).

[GR02] Gorochov, A. V., & A. P. Rasnitsyn. 2002. Superorder Gryllidea Laicharting, 1781 (=Orthopteroidea Handlirsch, 1903). In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 293–303. Kluwer Academic Publishers: Dordrecht.

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