Torymidae

 Female Torymoides kiesenwetteri, photographed by El Gritche.

Belongs within: Chalcidoidea.
Contains: Monodontomerini, Podagrionini, Torymini, Microdontomerini.

The Toryminae as recognised herein was supported as monophyletic in the molecular phylogenetic analysis of Chalcidoidea by Heraty et al. (2013); it had been treated as two separate subfamilies Toryminae and Monodontomerinae by Bouček (1988). The majority of members of this clade are parasitic; many are metallic in coloration. Members of the Podagrionini and Chrysochalcissini have enlarged hind femora with a number of ventral teeth; a single ventral tooth may be present on a less enlarged hind femur in Monodontomerini and Microdontomerini. The Podagrionini are parasites of mantis oothecae (Bouček 1988).

<==Torymidae [Callimonidae]HB13
    |--ThaumatoryminaeB88
    `--Toryminae [Monodontomerinae, Torymoidini]HB13
         |  i. s.: Odopoia Walker 1871 (see below for synonymy)B88
         |           |--*O. atraB88
         |           |--O. dentatinota (Girault 1925) [=*Australtorymus dentatinotus; incl. *Ua maria Girault 1929]B88
         |           |--O. josephinae Bouček 1988B88
         |           `--O. philippiae [=*Pauliana philippiae]B88
         |--Chalcimerus [Chalcimerini]B88
         |    `--C. borceaiHB13
         |--+--Platykula albihirtaMH11
         |  `--Boucekinus tatianaeMH11
         `--+--Glyphomerus stigmaHB13
            `--+--+--MonodontomeriniHB13
               |  `--+--PodagrioniniHB13
               |     `--PalachiiniHB13
               |          |--Palachia hayatiMH11
               |          `--Propalachia borneanaHB13
               `--+--+--ToryminiHB13
                  |  `--PseudotorymusHB13
                  |       |--P. lazulellusGZ96
                  |       `--P. sapphyrinusHB13
                  `--+--MicrodontomeriniHB13
                     `--Echthrodape Burks 1969HB13, B88 [Echthrodapinae]
                          |--*E. africanaB88
                          `--E. papuana Bouček 1988B88
Torymidae incertae sedis:
  Eotorymites Hong 2002H02
    `--*E. chloribrunneus Hong 2002H02
  Huaxiatorymus Hong 2002H02
    `--*H. castaneus Hong 2002H02
  Leptotorymites Hong 2002H02
    `--*L. xilutianensis Hong 2002H02
  Eomonodontomerites Hong 2002H02
    `--*E. azulichloriticus Hong 2002H02
  IdiomacromerusG05
    |--I. silybi Askew 2004F05
    `--I. urospermi Askew 2004F05
  Palaeotorymus renzii (Peñalver & Engel 2006) [=Perilampus renzii]HD09
  Neopalachia bouceki Grissell 1980P92
  Zophodetus Grissell 1980P92
    `--*Z. woodruffi Grissell 1980P92

Odopoia Walker 1871 [incl. Australtorymus Girault 1925, Pauliana Risbec 1952, Ua Girault 1929; Odopoiini]B88

*Type species of generic name indicated

References

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[F05] Fernández, J. 2005. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (IX). Graellsia 61 (2): 261–282.

[G05] Grissell, E. E. 2005. A review of North American species of Microdontomerus Crawford (Torymidae: Hymenoptera). Journal of Hymenoptera Research 14 (1): 22–65.

[GZ96] Grissell, E. E., & R. S. Zack. 1996. Torymidae (Hymenoptera) new to Washington State. Proceedings of the Entomological Society of Washington 98 (4): 827–828.

[HB13] Heraty, J. M., R. A. Burks, A. Cruaud, G. A. P. Gibson, J. Liljeblad, J. Munro, J.-Y. Rasplus, G. Delvare, P. Janšta, A. Gumovsky, J. Huber, J. B. Woolley, L. Krogmann, S. Heydon, A. Polaszek, S. Schmidt, D. C. Darling, M. W. Gates, J. Mottern, E. Murray, A. D. Molin, S. Triapitsyn, H. Baur, J. D. Pinto, S. van Noort, J. George & M. Yoder. 2013. A phylogenetic analysis of the megadiverse Chalcidoidea (Hymenoptera). Cladistics 29: 466–542.

[HD09] Heraty, J. M., & D. C. Darling. 2009. Fossil Eucharitidae and Perilampidae (Hymenoptera: Chalcidoidea). Zootaxa 2306: 1–16.

[H02] Hong Y. 2002. Amber Insect of China. Beijing Scientific and Technological Publishing House.

[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PloS One 6 (11): e27023.

[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.

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