Male (upper) and female Torymus bedeguaris, copyright Josef Dvořák.

Belongs within: Torymidae.

The Torymini are a group of torymid wasps in which the groove between the mesopleuron and metapleuron is straight or slightly curved, and the gaster is sessile or on a shiny, usually pale petiole. Most species are parasitoids of gall-forming or wood-boring insects; some are phytophagous seed-eaters (Bouček 1988).

In most genera, the gaster is sessile with the ovipositor being at most a little longer than the body in females. The genus Ecdamua has a pale, shiny petiole and the ovipositor is more than three times the length of the body. The Australian Ovidia conicicollis is a distinctively long, slender-bodied species with a conical pronotum, a lamellate scapus in the female that exceeds the vertex in height, and a long ovipositor covered in rough hairs (Bouček 1988).

    |--Ecdamua Walker 1862MH11, B88 [incl. Monodontomerella Girault 1915B88]
    |    |--*E. macrotelusB88
    |    |--E. indica [incl. Amonodontomerus indicus Ahmad 1946 non Ecdamua indica]B88
    |    `--E. longipilum (Girault 1925) [=*Monodontomerella longipilum]B88
    `--+--Physothorax bidentulusMH11
       `--Torymus Dalman 1820MH11, B88 (see below for synonymy)
            |--*T. bedeguaris (Linnaeus 1758)B88, L58 [=Ichneumon bedeguarisB88, *Callimome bedeguarisB88, Cynips bedeguarisL02]
            |--T. aeneoscapusGZ96
            |--T. albomaculatus Lucas 1849E12
            |--‘Syntomaspis’ apicalisA71
            |--T. aterA83
            |--T. auratusA71
            |--T. bicoloratusGZ96
            |--‘Callimome’ caburus Walker 1839W39
            |--T. capillaceusGZ96
            |    |--T. c. capillaceusGZ96
            |    `--T. c. albitarsisGZ96
            |--T. cecidomyiaeGZ96
            |--T. chrysochlorusGZ96
            |--T. cingulatusA71
            |--T. citripesGZ96
            |--T. coloradensisGZ96
            |--‘Syntomaspis’ cyaneaA71
            |--T. druparum Boheman 1834B88 [=Syntomaspis druparumRD77]
            |--T. eurynotus [=*Syntomaspis eurynotus]B88
            |--T. fagopirumGZ96
            |--T. josephinae Bouček 1988B88
            |--T. kinseyiGZ96
            |--T. koebeleiGZ96
            |--T. longistigmusGZ96
            |--T. nigricornisA71
            |--T. propinquusRD77
            |--T. solitariusGZ96
            |--‘Callimome’ sulcius Walker 1839W39
            |--T. thalassinusGZ96
            |--T. varians (Walker 1833)B88 [=Callimome variansB88, Syntomaspis variansR70]
            `--T. ventralisMS01
Torymini incertae sedis:
  Ovidia Girault 1924B88
    `--*O. conicicollis Girault 1924B88
  Austorymus Bouček 1988B88
    `--*A. nitidus Bouček 1988B88
  Mesodiomorus Strand 1911B88
    `--*M. compressusB88
  Diomorus Walker 1834B88
    |--*D. nobilisB88
    |--D. armatus (Boheman 1834) [=Torymus armatus]B88
    |--D. calcaratusBM76
    |--D. orientalis Masi 1926 [incl. Macrodontomerus silvifilia Girault 1927, D. silvifilia]B88
    `--D. zabriskiiGZ96

Torymus Dalman 1820MH11, B88 [=Callimome Spinola 1811 (nom. rej.)B88; incl. Misocampe Latreille 1817B88, Misocampus Dufour 1847B88, Syntomaspis Förster 1856B88]

*Type species of generic name indicated


[A83] Adolph, E. 1883. Zur Morphologie der Hymenopterenflügel. Zugleich ein Beitrag zu den Fragen der Speciesbildung und des Atavismus. Verhandlungen der Kaiserlichen Leopoldinisch-Carolinischen Deutschen Akademie der Naturforscher [Nova Acta Academiae Caesareae Leopoldino-Carolinae Germanicae Naturae Curiosorum] 46 (2): 41–132, pls 1–6.

[A71] Askew, R. R. 1971. Parasitic Insects. Heinemann Educational Books: London.

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[GZ96] Grissell, E. E., & R. S. Zack. 1996. Torymidae (Hymenoptera) new to Washington State. Proceedings of the Entomological Society of Washington 98 (4): 827–828.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.

[MS01] Mocsáry, A., & V. Szépligeti. 1901. Hymenopterák [Hymenopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 121–169. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PloS One 6 (11): e27023.

[RD77] Richards, O. W., & R. G. Davies. 1977. Imms’ General Textbook of Entomology 10th ed. vol. 2. Classification and Biology. Chapman and Hall: London.

[R70] Riek, E. F. 1970. Hymenoptera (wasps, bees, ants). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 867–959. Melbourne University Press.

[W39] Walker, F. 1839. Monographia Chalciditum vol. 2. Species collected by C. Darwin, Esq. Hyppolite Baillière: London.

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