Trechus quadristriatus, copyright Nikolai Vladimirov.

Belongs within: Trechinae.

The Trechini are typically small carabid beetles with broad, curved frontal grooves on the head (Roig-Juñent & Cicchino 2001).

Characters (from Roig-Juñent & Cicchino 2001): Head dorsally with frontal grooves curved, expanded to genae and ventral surface, at middle distance between eye and adjacent groove subequal to distance between grooves; two supraorbital setae on each side; maxillary palpomere 3 not densely pubescent; terminal maxillary and labial palpomeres not much smaller than penultimate palpomeres; glossal sclerite with six or more setae; antennal cleaner anisochaetus and setal band that is divided into large distal region and proximal cleaning arc, anterior row of setae absent, channel deep, short, median expansion well-developed anteriorly, tibia compressed anterioposteriorly; anterior tarsi with squamosetae in single row on long axis, with shafts distinct dorsoventrally, flattened distally, and laterally extended into almost rectangular parallel plates.

    |--Kenodactylus [Aepina]R-JC01
    |    `--K. audouiniR-JC01
    `--+--Aemalodera [Homaloderina]R-JC01
       |    `--A. centromaculataR-JC01
            |--Trechisibus antarcticusR-JC01
            `--Trechus Clairville 1806BB01
                 |  i. s.: T. amplicollis Fairmaire 1859OT05
                 |         T. angusticollisOT05
                 |         T. apoduvalipenis Salgado & Ortuño 1998OT05
                 |         T. austriacusOT05
                 |         T. barnevillei Pandellé 1867OT05
                 |         T. collarisR26
                 |         T. distigmaOT05
                 |         T. escalerai Abeille 1903OT05
                 |         T. fulvusR26
                 |         T. marcilhaci Pham 1987OT05
                 |         T. pandelleiOT05
                 |         T. pieltaini Jeannel 1920OT05
                 |         T. pisuenensis Ortuño & Toribio 2005F05
                 |         T. pyrenaeusOT05
                 |         T. saxicola Putzeys 1870OT05
                 |           |--T. s. saxicolaOT05
                 |           `--T. s. desbordesi Gaudin in Colas & Gaudin 1935OT05
                 |         T. tingitanusOT05
                 |         T. uhagoniOT05
                 |--T. (Trechus)BB01
                 |    |--T. (T.) apicalis Motschulsky 1845BB01
                 |    |--T. (T.) obtusus Erichson 1837BB01
                 |    |--T. (T.) quadristriatus (Schrank 1781)BB01
                 |    `--T. (T.) rubens (Fabricius 1801)BB01 [=Harpalus (T.) rubensG20]
                 `--T. (Microtrechus Jeannel 1927)BB01
Trechini incertae sedis:
  Paradivalius netolitzkyi Müller 1909PN02
  Neotrechus dalmatinus (Miller 1861)PN02
  Trechoblemus seriesBB01
    |--Pseudanophthalmus Jeannel 1920 [incl. Aphanotrechus Barr 1960]BB01
    |--Nelsonites Casale & Laneyrie 1982BB01
    |--Blemus Dejean 1821 [incl. Lasiotrechus Ganglbauer 1892]BB01
    |    `--B. discus (Fabricius 1792)BB01 [=Lasiotrechus discusB90]
    |--Neaphaenops Jeannel 1920BB01
    |    `--N. tellkampfi (Erichson 1844)BB01
    `--Trechoblemus Ganglbauer 1892BB01
         |--T. micros (Herbst 1784)BB01
         `--T. westcotti Barr 1972BB01
  Trechimorphus Jeannel 1927B74, LM87
    |--*T. diemenensis (Bates 1878) [=Trechus diemenensis]LM87
    |--T. apterus Moore 1972LM87
    |--T. brunneus Moore 1972LM87
    |--T. semipunctatus Moore 1972LM87
    |--T. solidior (Blackburn 1901) [=Trechus solidior]LM87
    `--T. westraliensis Moore 1972LM87
  Eutrechus Moore 1972B74, LM87
    |--*E. otwayensis (Moore 1960) [=Sloanella otwayensis]LM87
    |--E. barringtonensis Moore 1972LM87
    |--E. coxi (Sloane 1911) [=Trechus coxi]LM87
    `--E. gippslandicus (Sloane 1923) [=Trechus gippslandicus]LM87
  Darlingtonea seriesBB01
    |--Darlingtonea Valentine 1952BB01
    |    `--D. kentuckensis Valentine 1952BB01
    `--Ameroduvalius Valentine 1952BB01
         `--A. jeanneli Valentine 1952BB01
  Xenotrechus Barr & Krekeler 1967BB01
  Anophthalmus Sturm 1844BB01

*Type species of generic name indicated


[BB01] Ball, G. E., & Y. Bousquet. 2001. Carabidae Latreille, 1810. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 32–132. CRC Press: Boca Raton.

[B90] Bell, R. T. 1990. Insecta: Coleoptera Carabidae adults and larvae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1053–1092. John Wiley & Sones: New York.

[B74] Britton, E. B. 1974. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers. Supplement 1974 pp. 62–89. Melbourne University Press.

[F05] Fernández, J. 2005. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (IX). Graellsia 61 (2): 261–282.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[OT05] Ortuño, V. M., & M. Toribio. 2005. Descripción de un nuevo Trechus Clairville, 1806 (Coleoptera, Carabidae, Trechini) de los montes cantábricos orientales (norte de España). Graellsia 61 (1): 115–121.

[PN02] Poggi, R., G. Nonveiller, A. Colla, D. Pavićević & T. Rađa. 2001–2002. Thaumastocephalini, a new tribe of Pselaphinae for Thaumastocephalus folliculipalpus n. gen., n. sp., an interesting new troglobious species from central Dalmatia (Croatia) (Coleoptera, Staphylinidae, Pselaphinae). Annali del Museo Civico di Storia Naturale “Giacomo Doria” 94: 1–20.

[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.

[R-JC01] Roig-Juñent, S., & A. C. Cicchino. 2001. Chaltenia patagonica, new genus and species belonging to Chalteniina, a new subtribe of Zolini (Coleoptera: Carabidae). Canadian Entomologist 133: 651–670.

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