Trhypochthoniidae

Dorsal (left) and ventral views of Allonothrus neotropicus, from Balogh & Balogh (1992).

Belongs within: Holonota.
Contains: Trhypochthonius.

Allonothrus is a genus of tropical and subtropical oribatid mites bearing elongate setae on the posterior part of the notogaster.

<==Trhypochthoniidae [Trhypochthoniellidae, Trhypochthoniinae]
    |  i. s.: Afronothrus Wallwork 1961S04
    |           |--*A. incisivus Wallwork 1961S04 (see below for synonymy)
    |           `--A. arboreus Ramani & Haq 1992S04
    |         Mainothrus Choi 1996 [incl. Altrhypochthonius Weigmann 1997]S04
    |           |--*M. aquaticus Choi 1996S04
    |           |--M. badius (Berlese 1905) [=Trhypochthonius badius]S04
    |           `--M. breviclava (Hammer 1958) [=Trhypochthonius breviclava]S04
    |         Palaeochthonius krasilovi Krivolutsky & Ryabinin 1976S93
    |         Juracarus serratus Krivolutsky & Ryabinin 1976S93
    |--Allonothrus Hammen 1953DW10, S04 [Allonothridae, Parallonothridae]
    |    |--A. (Allonothrus)S04
    |    |    |--*A. (A.) schuilingi Hammen 1953S04
    |    |    |--A. (A.) foveolatus Pérez-Íñigo & Baggio 1988S04
    |    |    |--A. (A.) giganticus Haq 1978S04
    |    |    |--A. (A.) indicus Bhaduri & Raychaudhuri 1968S04
    |    |    |--A. (A.) monensis Ghosh & Bhaduri 1978S04
    |    |    |--A. (A.) monodactylus Wallwork 1960S04
    |    |    |--A. (A.) neotropicus Balogh & Mahunka 1969S04
    |    |    |--A. (A.) pararusseolus Subías & Sarkar 1982S04
    |    |    |--A. (A.) pyriformis (Berlese 1913) [=Angelia pyriformis]S04
    |    |    |--A. (A.) russeolus Wallwork 1960S04
    |    |    |    |--A. r. russeolusS04
    |    |    |    `--A. r. reticulatus Hammer 1972S04
    |    |    |--A. (A.) sinicus Wang & Norton 1988S04
    |    |    `--A. (A.) tuxtlasensis Palacios-Vargas & Iglesias 1997S04
    |    `--A. (Pseudonothrus Balogh 1958)S04
    |         `--A. (*P.) hirtus (Balogh 1958)S04 (see below for synonymy)
    `--+--Trhypochthoniellus Willmann 1928SC19, S04 [incl. Hydronothrus Aoki 1964S04]
       |    |--T. longisetus (Berlese 1904) [=Trhypochthonius longisetus]S04
       |    |    |--T. l. longisetusS04 (see below for synonymy)
       |    |    |--T. l. brasilensis Pérez-Íñigo & Baggio 1988 [=T. setosus brasilensis]S04
       |    |    `--T. l. canadensis Hammer 1952 [=T. setosus canadensis]S04
       |    |--T. ashoroensis Fujikawa 2000S04
       |    |--T. badiusNK94
       |    |--T. crassus (Warburton & Pearce 1905)SC19, S04 (see below for synonymy)
       |    |--T. porticus Fujikawa 2000S04
       |    |--T. ramosus Hammer 1982S04
       |    `--T. reticulatus Willmann 1931S04
       `--+--Mucronothrus Trägardh 1931SC19, S04 [Mucronothridae]
          |    |--M. nasalis (Willmann 1929) [=Malaconothrus nasalis; incl. *Mu. rostratus Trägardh 1931]S04
          |    `--M. willmanni Norton, Behan-Pelletier & Wang 1996S04
          `--+--TrhypochthoniusDW10
             `--Archegozetes Grandjean 1931DW10, S04
                  |--*A. magnus (Sellnick 1925) [=Epilohmannia magna]S04
                  |    |--A. m. magnus (see below for synonymy)S04
                  |    `--A. m. mediosetosus Mahunka 1978S04
                  `--A. longisetosus Aoki 1965S04 (see below for synonymy)

*Afronothrus incisivus Wallwork 1961S04 [incl. A. incisivus maheensis Mahunka 1978S04, A. incisivus neotropicus Balogh & Mahunka 1974S04, A. incisivus paraguayensis Mahunka 1984S04, A. sulcatus Hammer 1972S04, A. incisivus sulcatusCH98]

Allonothrus (*Pseudonothrus) hirtus (Balogh 1958)S04 [=*Pseudonothrus hirtusS04; incl. A. ghanensis Wallwork 1961S04, P. ghanensisBB92]

Archegozetes longisetosus Aoki 1965S04 [=A. magnus longisetosusM78; incl. A. chamelensis Palacios-Vargas & Iglesias 1997S04, A. magnus indicus Bhaduri & Raychaudhuri 1968S04]

*Archegozetes magnus magnus (Sellnick 1925) [incl. A. tuberculatus Sarkar 1985, A. vercruzensis Palacios-Vargas & Iglesias 1997]S04

Trhypochthoniellus crassus (Warburton & Pearce 1905)SC19, S04 [=Nothrus crassusS04, Malaconothrus crassusCH98, Trimalaconothrus crassusCH98, Trhypochthonius crassusL87]

Trhypochthoniellus longisetus longisetus (Berlese 1904)S04 [incl. Hydronothrus aquariorum Fain & Lambrechts 1987S04, *Hydronothrus crispus Aoki 1964CH98, Trhypochthoniellus crispusSN98, Camisia excavata Willmann 1919S04, Trhypochthoniellus excavatusG39, Trhypochthonius excavataA70, Trhypochthonius excavatusCH98, Hydronothrus puniceus Habeeb 1981S04, Trhypochthonius (*Trhypochthoniellus) setosus Willmann 1928S04, Trhypochthonius trichosus Schweizer 1922S04, Tripochthonius (l. c.) trichosusS22]

*Type species of generic name indicated

References

[A70] Aoki, J. 1970. The oribatid mites of the islands of Tsushima. Bulletin of the National Science Museum, Tokyo 13 (3): 395–442.

[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 2. Plates. Hungarian Natural History Museum: Budapest.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[DW10] Dabert, M., W. Witalinski, A. Kazmierski, Z. Olszanowski & J. Dabert. 2010. Molecular phylogeny of acariform mites (Acari, Arachnida): strong conflict between phylogenetic signal and long-branch attraction artifacts. Molecular Phylogenetics and Evolution 56 (1): 222–241.

[G39] Grandjean, F. 1939. Observations sur les oribates (12e série). Bulletin du Muséum National d’Histoire Naturelle, 2e Série 11 (3): 300–307.

[L87] Luxton, M. 1987. The oribatid mites (Acari: Cryptostigmata) of J. E. Hull. Journal of Natural History 21: 1273–1291.

[M78] Mahunka, S. 1978. Neue und interessante Milben aus dem Genfer Museum XXVII. A first survey of the oribatid (Acari) fauna of Mauritius, Reunion and the Seychelles I. Revue Suisse de Zoologie 85 (1): 177–236.

[NK94] Norton, R. A., & J. B. Kethley. 1994. Ecdysial cleavage lines of acariform mites (Arachnida, Acari). Zoologica Scripta 23 (3): 175–191.

[SC19] Schaefer, I., & T. Caruso. 2019. Oribatid mites show that soil food web complexity and close aboveground-belowground linkages emerged in the early Paleozoic. Communications Biology 2: 387.

[S22] Schweizer, J. 1922. Beitrag zur Kenntnis der terrestrischen Milbenfauna der Schweiz. Verhandl. Naturf. Ges. Basel 33: 23–112, 4 pls.

[S93] Selden, P. A. 1993. Arthropoda (Aglaspidida, Pycnogonida and Chelicerata). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 297–320. Chapman & Hall: London.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

[SN98] Swift, S. F., & R. A. Norton. 1998. Preliminary report on oribatid mite (Acari: Oribatida) diversity in the Hawaiian Islands. Bishop Museum Occasional Papers 57: 1–44.

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