Red flour beetle Tribolium castaneum, copyright Udo Schmidt.

Belongs within: Tenebrionidae.

The Triboliini are small, parallel-sided tenebrionid beetles with reniform eyes (Matthews & Bouchard 2008).

Characters (from Matthews & Bouchard 2008): Oblong, parallel-sided, moderately convex to subcylindrical, glabrous, fuscous. Total length 2–6 mm. Edge of clypeus nearly straight in middle or shallowly emarginate. Eyes reniform. Labrum transverse, medial tormal arms of epipharynx either transverse, parallel to basal edge of labrum and mesally expanded into short anterior and posterior extensions, or strongly angled basad and not mesally expanded, without epitorma, setal pattern more or less symmetrical. Antennae short, not reaching base of prothorax, segments gradually or (usually) abruptly enlarging distally to form club, with simple setiform sensilla. Corpotentorium more or less arcuate. Mandibles with long scissorial lobe and small mola, apically bidentate, molar surface with or without fine striations. Maxilla with lacinial uncus in form of slender bristle-like tooth, palpi with apical segment fusiform. Prosternum long before coxae. Procoxal cavities open or closed internally. Mesocoxal cavities closed by mesepimeron. Metendosternite with large laminae, with short to moderate stem. Elytra with nine striae and scutellary striole, epipleura complete to apex. Wings present, venation unmodified, without medial or apical flecks, apical membrane about 40% of wing length. Legs not fossorial, subapical tarsomeres somewhat cupuliform. Plantar surfaces sparsely setose. Abdomen with defensive gland reservoirs present, large and with constricted exit ducts, without lateral stiffening prongs, without common volume. Ovipositor very variable, either with coxite lobe 4 strongly sclerotised and pointed, apically spatulate, or coxites wholly membranous without discernible lobation, gonostyles reduced and lateral or dorsal in position, coxite baculi transverse, paraproct baculi either longitudinal or oblique. Female tract without bursa, with single curved or coiled spermathecal tubule emerging from vagina separately from accessory gland, vaginal sclerite present or absent. Aedeagus with basal piece longer than apical.

<==Triboliini [Triboliina]
    |--Mycotrogus Horn 1870AT02
    |--Latheticus Waterhouse 1880MB08
    |    `--*L. oryzae Waterhouse 1880MB08
    |--Hypogena Dejean 1834 [incl. Ulosonia Laporte 1840]AT02
    |    `--‘Ulosonia’ tricorniAT02
    |--Tharsus LeConte 1862AT02
    |    `--T. seditiosus LeConte 1862AT02
    |--Lyphia Mulsant & Rey 1859 [incl. Lindia Blackburn 1888 non Dujardin 1841]MB08
    |    |--*L. ficicola Mulsant & Rey 1859MB08
    |    |--L. australis Gebien 1911 [=*Lindia angusta Blackburn 1888 (preoc.)]MB08
    |    |--L. grandis Carter 1926MB08
    |    `--L. tasmanica Champion 1894MB08
    `--Tribolium Macleay 1825MB08 (see below for synonymy)
         |--*T. castaneum (Herbst 1797) [=Colydium castaneum; incl. Dermestes navalis Fabricius 1775 (nom. rej.)]MB08
         |--T. antennatum Hinton 1948MB08
         |--T. apiculum Neboiss 1962MB08
         |--T. confusum Jacquelin du Val 1862MB08
         |--T. destructorB14
         |--T. ferrugineum Fab. 1781 [incl. T. cinnamomeum, T. testaceum]FS90
         |--T. madensC68
         |--T. myrmecophilum Lea 1904 [=T. (*Leanum) myrmecophilum]MB08
         `--T. waterhousei Hinton 1948MB08

Tribolium Macleay 1825MB08 [incl. Aphanotus LeConte 1862AT02, Eusemostene Gebien 1940AT02, Leanum Uyttenboogaart 1934MB08, Margus Dejean 1833AT02, Stene Stephens 1832AT02, Tenebrioloma Gebien 1910AT02]

*Type species of generic name indicated


[AT02] Aalbu, R. L., C. A. Triplehorn, J. M. Campbell, K. W. Brown, R. E. Somerby & D. B. Thomas. 2002. Tenebrionidae Latreille 1802. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 463–509. CRC Press.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).

[C68] Crowson, R. A. 1968. Concerning immunotaxonomy of Coleoptera. Systematic Zoology 17 (1): 95–97.

[FS90] Fleutiaux, E., & A. Sallé. 1890. Liste des coléoptères de la Guadeloupe et descriptions d’espèces nouvelles. Annales de la Société Entomologique de France, 6e série 9: 351–484.

[MB08] Matthews, E. G., & P. Bouchard. 2008. Tenebrionid Beetles of Australia: Descriptions of tribes, keys to genera, catalogue of species. Australian Biological Resources Study: Canberra.

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