Spinopilar moria, copyright BioEspeleo Consultoria Ambiental.

Belongs within: Gonyleptidae.

The Tricommatinae are a group of small short-legged harvestmen found in southern Brazil and Paraguay. The group is characterised by an elongate coxa II which widely surpasses coxa III in ventral view (Kury 2002).

<==Tricommatinae [Olynthoidae, Triacommatinae, Trichommantinae, Trichommatinae, Tricommatidae]
    |--Caramaschia Kury 2002K02
    |    `--*C. singularis Kury 2002K02
    |--Pseudophalangodes Roewer 1912K02 [=Pseudophalangodus (l. c.)K03]
    |    `--*P. unicolor Roewer 1912K03
    |--Rivetinus Roewer 1919R23, K03 [=Ruvetinus (l. c.)K03]
    |    `--*R. minutus Roewer 1919K03
    |--Neoarminda Özdikmen & Kury 2006 [=Arminda Roewer 1949 non Krauss 1892]OK06
    |    `--*N. colatinae (Soares & Soares 1946) [=Phalangodella colatinae, *Arminda colatinae]OK06
    |--Bebedoura Roewer 1949K03
    |    `--*B. rugosa Roewer 1949K03
    |--Bissulla Roewer 1929K03 [=Bissula (l. c.)KA-Z11]
    |    `--*B. paradoxa Roewer 1929K03
    |--Bunostigma Mello-Leitão 1935K03
    |    `--*B. singulare Mello-Leitão 1935K03, KA-Z11, K03 [=B. singularisK03]
    |--Lanesoares Roewer 1949K03
    |    `--*L. inermis (Soares 1944) [=Phalangodella inermis]K03
    |--Otuquisa Roewer 1927K03
    |    `--*O. distincta Roewer 1927K03
    |--Pararezendesius Soares 1972K03
    |    `--*P. luridus Soares 1972K03
    |--Poecilosophus Mello-Leitão 1949K03, KA-Z11 [=Soaresula Roewer 1949K03]
    |    `--*P. melloleitaoi (Soares 1945) [=Paratricommatus melloleitaoi, *Soaresula melloleitaoi]K03
    |--Rezendezius Soares 1945K03
    |    `--*R. lanei Soares 1945K03
    |--Saladonus Roewer 1927K03
    |    `--*S. singularis Roewer 1927K03
    |--Segundolus Roewer 1949K03
    |    `--*S. insignitus Roewer 1949K03
    |--Taquara Mello-Leitão 1936 [=Taguara (l. c.); incl. Therezopachylus Soares 1972]K03
    |    `--*T. pilosa Mello-Leitão 1936 [incl. *Therezopachylus fuscus Soares 1972]K03
    |--Berlesecaptus Mello-Leitão 1940KP-R07, K03
    |    `--*B. convexus Mello-Leitão 1940K03
    |--Heteromeloleptes Mello-Leitão 1931K02 [incl. Derzius Roewer 1949K03, Melloceras Roewer 1949K03, Mellops Roewer 1949K03]
    |    |--*H. padbergi Mello-Leitão 1931 (see below for synonymy)K03
    |    `--H. fasciatus (Mello-Leitão 1932) [=Phalangodella fasciata, *Mellops fasciatus]K03
    |--Bresslauius Mello-Leitão 1935 [incl. Friburguesia Soares 1946, Pinocchio Mello-Leitão 1940]K03
    |    |--*B. hirsutus Mello-Leitão 1935 (see below for synonymy)K03
    |    `--B. debilis (Mello-Leitão 1940) [=*Pinocchio debilis]K03
    |--Machairoscelis Kury 2003K03
    |    |--*M. armatipes (Soares 1972) [=Corcovadesia armatipes]K03
    |    `--M. singularis (Soares & Soares 1949) [=Paramitraceras singularis]K03
    |--Paratricommatus Piza 1943K03
    |    |--*P. modestus Piza 1943K03
    |    `--P. mahnerti Soares & Soares 1985K03
    |--Tricommatus Roewer 1912K02
    |    |--*T. brasiliensis Roewer 1912K02
    |    |--T. fulvus Soares 1966 [incl. Prosontes brasiliensis Soares 1966 non T. brasiliensis Roewer 1912]CK20
    |    `--T. giuponii (Kury 2003)A19
    |--Simonoleptes Soares & Soares 1954K02
    |    |--*S. latifemur Soares & Soares 1954K03
    |    |--S. insulanus Soares & Soares 1966K03
    |    `--S. obstectispiracula Spares & Soares 1954K03
    |--Arruda Mello-Leitão 1940K03
    |    |--*A. insignis Mello-Leitão 1940K03
    |    |--A. mutilata Mello-Leitão 1940K03
    |    `--A. pectinata Mello-Leitão 1940K03
    |--Corcovadesia Soares & Soares 1954K03, KA-Z11 (see below for synonymy)
    |    |--*C. inermis Soares & Soares 1954 [=Liops inermis]K03
    |    |--C. hexabunus (Mello-Leitão 1940)KA-Z11 [=*Liops hexabunusK03]
    |    `--C. venefica (Soares 1966)KA-Z11 [=*Buzioleptes veneficusK03, Liops veneficusK03]
    `--Camarana Mello-Leitão 1935K02
         |--*C. minor Mello-Leitão 1935K03
         |--C. bicoloripes Soares 1974K03
         |--C. flavipalpi Soares 1945K03
         |--C. rousseti Soares & Soares 1946K03
         `--C. unica Soares 1944K03

*Bresslauius hirsutus Mello-Leitão 1935 [=Pucrolia hirsuta; incl. *Friburguesia hirsuta Soares 1946 non Bresslauius hirsutus Mello-Leitão 1935]K03

Corcovadesia Soares & Soares 1954K03, KA-Z11 [incl. Buzioleptes Soares 1966K03, Liops Mello-Leitão 1940 nec Fieber 1870 nec Gidley 1906KA-Z11]

*Heteromeloleptes padbergi Mello-Leitão 1931 [incl. Pseudopachylus anomalus Mello-Leitão 1935, *Derzius anomalus, Pseudophalangodes bimaculatus Mello-Leitão 1932, *Melloceras bimaculatus, Phalangodella jandirae Soares & Soares 1970]K03

*Type species of generic name indicated


[A19] Acosta, L. E. 2019. A relictual troglomorphic harvestman discovered in a volcanic cave of western Argentina: Otilioleptes marcelae, new genus, new species, and Otilioleptidae, new family (Arachnida, Opiliones, Gonyleptoidea). PLoS One 14 (10): e0223828.

[CK20] Caramori, L., & A. B. Kury. 2020. On a tricommatine from Paraná described in two different families (Laniatores: Gonyleptidae). In: Kury, A. B., A. C. Mendes, L. Cardoso, M. S. Kury & A. A. Granado. 2020. WCO-Lite: Online world catalogue of harvestmen (Arachnida, Opiliones). Version 1.0—Checklist of all valid nomina in Opiliones with authors and dates of publication up to 2018 p. 48. Published by the author: Rio de Janeiro.

[K02] Kury, A. B. 2002. A new genus of Tricommatinae from eastern Brazil (Opiliones Laniatores Gonyleptidae). Tropical Zoology 15: 209–218.

[K03] Kury, A. B. 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Ibérica de Aracnología, volumen especial monográfico 1: 1–337.

[KA-Z11] Kury, A. B., & M. A. Alonso-Zarazaga. 2011. Addenda and corrigenda to the “Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones)”. Zootaxa 3034: 47–68.

[KP-R07] Kury, A. B., & R. Pinto-da-Rocha. 2007. Gonyleptidae Sundevall, 1833. In: Pinto-da-Rocha, R., G. Machado & G. Giribet (eds) Harvestmen: The Biology of Opiliones pp. 196–203. Harvard University Press: Cambridge (Massachusetts).

[OK06] Özdikmen, H., & A. B. Kury. 2006. Three homonymous generic names in Araneae and Opiliones. Journal of Arachnology 34 (1): 279–280.

[R23] Roewer, C.-F. 1923. Die Weberknechte der Erde: Systematische Bearbeitung der bisher bekannnten Opiliones. Gustav Fischer: Jena.

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