Dorsum of Trionyxella granulata, from Roewer (1935).

Belongs within: Assamiidae.

The Trionyxellinae are a group of harvestmen found in southern Africa and Asia, characterised by the presence of a pseudonychium between the true claws.

Characters (from Roewer 1935): Anterior margin of the carapace unarmed medially; spiracles concealed by chitinous projections; femur of pedipalp with short ventral teeth; distitarsus I two-segmented; tarsi III and IV with simple claws, with pseudonychium present over claws.

<==Trionyxellinae [Trionyxellidae]
|–Triaenopodium Roewer 1915 R35
|    `–T. tarsalis Roewer 1915 R35
|–Kandyca Roewer 1915 R35
|    `–K. minima Roewer 1915 R35
|–Nuwaria Roewer 1915 R35
|    `–N. granulata Roewer 1915 R35
|–Nilgirius Roewer 1915 R35
|    `–N. scaber Roewer 1915 R35
|–Calloristus Roewer 1935 R35
|    `–*C. granipes Roewer 1935 R35
|–Balnissa Roewer 1935 R35
|    `–*B. parva Roewer 1935 R35
|–Brysma Roewer 1935 R35
|    `–*B. atra Roewer 1935 R35
|–Trionychiperna Roewer 1929 R35
|    `–T. carli Roewer 1929 R35
|–Sampitius Roewer 1935 R35
|    `–*S. opacus Roewer 1935 R35
|–Dulitellus Roewer 1935 R35
|    |–*D. maculatus Roewer 1935 R35
|    `–D. sarawakensis WP-G10
|–Namutonia Lawrence 1931 R35 [incl. Gobabisia Roewer 1940 S92]
|    |–*N. scabra Lawrence 1931 S92
|    `–N. wuehlischi (Roewer 1940) [=Gobabisia wuehlischi] S92
|–Trionyxella Roewer 1912 R35
|    |–T. clavipus Roewer 1912 R35
|    `–T. granulata Roewer 1935 R35
`–Randilella Lawrence 1963 L63
`–*R. transvaalensis Lawrence 1963 S92

*Type species of generic name indicated


[L63] Lawrence, R. F. 1963. The Opiliones of the Transvaal. Annals of the Transvaal Museum 24 (4): 275–304.

[R35] Roewer, C.-F. 1935. Alte und neue Assamiidae. Weitere Weberknechte VIII. (8. Ergänzung der “Weberknechte der Erde” 1923). Veröffentlichungen aus dem Deutschen Kolonial- und Uebersee-Museum in Bremen 1: 1–168.

[S92] Staręga, W. 1992. An annotated check-list of Afrotropical harvestmen, excluding the Phalangiidae (Opiliones). Annals of the Natal Museum 33 (2): 271–336.

[WP-G10] Willemart, R. H., A. Pérez-González, J.-P. Farine & P. Gnaspini. 2010. Sexually dimorphic tegumental gland openings in Laniatores (Arachnida, Opiliones), with new data on 23 species. Journal of Morphology 271 (6): 641–653.

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