Cerobasis guestfalica, photographed by Brian Valentine.

Belongs within: Psocodea.

The Trogioidea is a group of small barklice with wings reduced or absent. The majority of living species are included in the subfamily Trogiinae with the South American genus Anomocopeus sometimes treated as a distinct family. The Archaeatropidae and Empheriinae are known only from the fossil record.

Characters (from Li 2002; translated using Mandarin Tools, all errors my own): Small size, wings (if present) not more than buds. Antennae with more than twenty segments, not annulated. Epicranial sutures complete or lateral arms absent; eyes large and not hairy, ocelli absent or three present, closely placed; lacinia tip not toothed or with only one large tooth, labial palp with four slender segments, terminal segment axe-shaped. Tarsus three-segmented, claw without preapical tooth.

    |--Anomocopeus Badonnel 1967 [Anomocopeidae]L02
    `--Trogiidae [Atropidae]SN08
         |  i. s.: EolepinotusP92
         |--EmpheriaP92 [Empheriidae, EmpheriinaeNL07]
              |--Myrmicodipnella Enderlein 1909M93
              |    `--*M. aptera Enderlein 1909M93
              |--Helminotrogia Li 2002 [=Heliminotrogia]L03
              |    `--*H. bipunctata Li 2002 [=H. bipuctata, H. bipunctatia]L03
              |--Phlebotrogia Li 2002 [=Philebotrogia, Phlebotrogius]L03
              |    `--*P. chinensis Li 2002L03
              |--Trogium Illiger 1798SN08 [=Troctes Burmeister 1839B50; incl. Atropos Leach 1815L98, Clothilla Westwood 1841L98]
              |    |--*T. pulsatorium (Linnaeus 1758)SN08 (see below for synonymy)
              |    |--‘Troctes’ atroposR13
              |    `--‘Troctes’ divinatorius (Müll. 1776)E07
              |--Lepinotus Heyden 1850SN08 (see below for synonymy)
              |    |--*L. inquilinus Heyden 1850SN08 (see below for synonymy)
              |    |--L. angolensis Badonnel 1955B55
              |    |--L. huoni Schmidt & New 2008SN08
              |    |--L. patruelis Pearman 1931SN08 (see below for synonymy)
              |    |--L. reticulatus Enderlein 1905SN08
              |    |--L. stoneaeSN08
              |    |--L. tasmaniensis Hickman 1934SN08
              |    `--L. vermicularis Lienhard 1996L98
              `--Cerobasis Kolbe 1882SN08 (see below for synonymy)
                   |  i. s.: C. albipes Lienhard 1996L98
                   |         C. amorosa Lienhard 1995L98
                   |         C. denticulata Lienhard 1996L98
                   |         C. ericacea Baz 1993 [=C. ericaceus]L98
                   |         C. insularis Lienhard 1996L98
                   |         C. longicornis Baz 1993L98
                   |         C. nigra Lienhard 1996L98
                   |         C. pineticola Baz 1991L98
                   |         C. rosae Baz 1993L98
                   |         C. socotrae Lienhard 1995L98
                   |--Group IM93
                   |    |--C. guestfalica (Kolbe 1880)M93 (see below for synonymy)
                   |    |--C. alfredi Lienhard 1984M93, L98
                   |    |--C. annulata (Hagen 1865)M93 (see below for synonymy)
                   |    |--C. canariensis (Enderlein 1910)M93, L98 (see below for synonymy)
                   |    |--C. lapidaria Badonnel 1955M93, B55 [=C. lapidariusMG56]
                   |    |--C. maculicepsM93
                   |    |--C. maderensis Lienhard 1983M93, L98
                   |    `--C. rectaM93
                   `--Group IIM93
                        |--C. caboverdensis Lienhard 1984M93, L98
                        |--C. harteni Lienhard 1984M93, L98
                        |--C. intermedia Lienhard 1984M93, L98
                        |--C. lambdaM93
                        |--C. papillataM93
                        `--C. trepticaM93

Cerobasis Kolbe 1882SN08 [incl. Albardia Jakobson in Jakobson & Bianchi 1904 (preoc.)L98, Hyperetes Kolbe 1880 (preoc.)L98, Myopsocnema Enderlein 1905L98, Tichobia Kolbe 1882L98, Zlinia Obr 1948L98]

Cerobasis annulata (Hagen 1865)M93 [=Clothilla annulataNL07, Atropos annulataM93, *Myopsocnema annulataNL07; incl. Cerobasis bundyi Turner 1977L98, *Zlinia multispinosa Obr 1948NL07, L98, Ce. multispinosaM93]

Cerobasis canariensis (Enderlein 1910)M93, L98 [=Myopsocnema canariensisL98; incl. C. anagaensis Baz 1993L98, C. palmensis Baz 1993L98]

Cerobasis guestfalica (Kolbe 1880)M93 [=*Hyperetes guestfalicusNL07; incl. *Tichobia alternans Kolbe 1882NL07, L98, *Albardia alternansNL07, *Cerobasis muraria Kolbe 1882SN08, L98, Hyperetes pinicola Kolbe 1881L98, H. tessulatus Hagen 1883L98]

Lepinotus Heyden 1850SN08 [incl. Cuixa Navás 1927L98, Heterolepinotus Obr 1948L98, Paradoxenus Molshulsky 1852L98, Paradoxides Motschulsky 1851 (preoc.)L98]

*Lepinotus inquilinus Heyden 1850SN08 [=Clothilla inquilinaM93, Atropos inquilinaM93; incl. *Cuixa canaria Navás 1927NL07, L98, A. distincta Kolbe 1888L98, Clothilla distinctaM93, Cl. picea Hagen 1861L98, Atropos piceaM93, Lepinotus piceusM93, *Paradoxides psocoides Motschulsky 1851NL07, L98, *Paradoxenus psocoidesNL07, Termes pulsatorium Scopoli 1763 non Linnaeus 1758M93, Atropos sericea Kolbe 1883L98, Lepinotus sericeusM93, Paradoxenus tubericola Hagen 1858G74]

Lepinotus patruelis Pearman 1931SN08 [incl. *Heterolepinotus quadrispinosus Obr 1948NL07, L98, L. (Heterolepinotus) quadrispinosusNL07]

*Trogium pulsatorium (Linnaeus 1758)SN08 [=Termes pulsatoriumSN08, Atropos pulsatoriaM93, Clothilla pulsatoriaM93, Hemerobius pulsatorius Fabricius 1775M93, Psocus pulsatoriusM93, Psylla pulsatoria Billburg 1820M93, *Troctes pulsatoriusB50; incl. Termes lignarium De Geer 1778L98, *Atropos lignariaNL07, Clothilla ocelloria Weber 1906L98, *C. studiosa Westwood 1841NL07, L98]

*Type species of generic name indicated


[B55] Badonnel, A. 1955. Psocoptères de l’Angola. Diamang Publicações Culturais 26: 11–267.

[B50] Broadhead, E. 1950. A revision of the genus Liposcelis Motschulsky with notes on the position of this genus in the order Corrodentia and on the variability of ten Liposcelis species. Transactions of the Royal Entomological Society of London 101 (10): 335–388, pls 1–3.

[E07] Enderlein, G. 1907. Wissenschaftliche Ergebnisse der Schwedischen Zoologischen Expedition nach dem Kilimandjaro, dem Meru und den Umgebenden Massaisteppen Deutsch-Ostafrikas 1905–1906 vol. 15. Corrodentia pt 2. Copeognatha. Almqvist & Wiksells Boktryckeri-A. B.: Uppsala.

[G74] Günther, K. K. 1974. Staubläuse, Psocoptera. Die Tierwelt Deutschlands 61: 1–315.

[L02] Li F. 2002. Psocoptera of China vol. 1. Science Press: Beijing.

[L98] Lienhard, C. 1998. Faune de France. France et Régions Limitrophes. 83. Psocoptères Euro-Méditerranéens. Fédération Française des Sociétés de Sciences Naturelles: Paris.

[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[MG56] Mockford, E. L., & A. B. Gurney. 1956. A review of the psocids, or book-lice and bark-lice, of Texas (Psocoptera). Journal of the Washington Academy of Sciences 46 (11): 353–368.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.

[R02] Rasnitsyn, A. P. 2002. Superorder Psocidea Leach, 1815. In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 125–133. Kluwer Academic Publishers: Dordrecht.

[R13] Reuter, O. M. 1913. Lebensgewohnheiten und Instinkte der Insekten bis zum Erwachen der sozialen Instinkte. R. Friedländer & Sohn: Berlin.

[SN08] Schmidt, E. R., & T. R. New. 2008. The Psocoptera (Insecta) of Tasmania, Australia. Memoirs of Museum Victoria 65: 71–152.

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