Ulodidae

 Syrphetodes decoratus, photographed by S. E. Thorpe.

Belongs within: Tenebrionoidea.

The Ulodidae are a Gondwanan family of beetles, members of which were formerly classified with the Tenebrionidae or Zopheridae. Many species have a vestiture of scales and bear waxy encrustations. The tarsal formula is 4-4-4 in the genus Meryx but 5-5-4 in Ulodinae, which also have expanded pronotal margins. Ulodinae are known to feed on basidiomycete fruiting bodies whereas Meryx are found under bark of fungus-infested logs (Lawrence & Britton 1991).

Synapomorphies (from Leschen & Rhode 2002): Antennal insertions visible in dorsal view, eyes entire or not anteriorly emarginate, procoxal cavities closed, lateral mesocoxal cavities open with exposed mesotrochantins, aedeagus with a dorsal tegmen.

<==Ulodidae
|--Syrphetodes decoratusMW15, LR02
`--+--Sirrhas limbatusMW15
`--Meryx [Merycidae, Merycinae]MW15
|--M. aequalis Blackb. 1891M96
`--M. rugosaMW15 [=Colydium (Meryx) rugosumG20]

Ulodidae incertae sedis:
TrachyderasLR02
TrachyderastesLR02
ArthopusLR02
BrouniphylaxLR02
ExohadrusLR02
Waitomophylax Leschen & Rhode 2004 [=Archaeophylax Leschen & Rhode 2002 non Kimmins 1953]LR04
`--*W. worthyi (Leschen & Rhode 2002) [=*Archaeophylax worthyi]LR04
UlodinaeLB91
|--PhaennisLB91
|--NotocerastesLB91
|--UlodesLR02
| |--U. variicornis Hope 1842M86
| `--U. verrucosus Erichs. 1842M86
|--GanymeLB91
| |--G. howitti Pascoe 1869M86
| `--G. sapphira Newm. 1842M86
`--DipsaconiaLB91
|--D. australis Hope 1845M86
|--D. bakewelli Pascoe 1860M86
`--D. pyritosa Pascoe 1850M86

*Type species of generic name indicated

References

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[LR02] Leschen, R. A. B., & B. E. Rhode. 2002. A new genus and species of large extinct Ulodidae (Coleoptera) from New Zealand. New Zealand Entomologist 25: 57–64.

[LR04] Leschen, R. A. B., & B. E. Rhode. 2004. A replacement name for Archaeophylax Leschen and Rhode (Coleoptera: Ulodidae). New Zealand Entomologist 27: 125.

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part IV. Proceedings of the Linnean Society of New South Wales, series 2, 1 (2): 259–380.

[M96] Masters, G. 1896. Catalogue of the described Coleoptera of Australia. Supplement, part II. Dytiscidae, Gyrinidae, Hydrophyllidae, Staphylinidae, Pselaphidae, Paussidae, Silphidae, Scaphididae, Histeridae, Phalacridae, Nitidulidae, Trogositidae, Colydiidae, Cucujidae, Cryptophagidae, Lathridiidae, Mycetophagidae, Dermestidae, Byrrhidae, Parnidae, Heteroceridae. Proceedings of the Linnean Society of New South Wales 21 (Suppl.): 695–754.

[MW15] McKenna, D. D., A. L. Wild, K. Kanda, C. L. Bellamy, R. G. Beutel, M. S. Caterino, C. W. Farnum, D. C. Hawks, M. A. Ivie, M. L. Jameson, R. A. B. Leschen, A. E. Marvaldi, J. V. McHugh, A. F. Newton, J. A. Robertson, M. K. Thayer, M. F. Whiting, J. F. Lawrence, A. Ślipiński, D. R. Maddison & B. D. Farrell. 2015. The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology 40 (4): 835–880.

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