Ulomini

Uloma culinaris, copyright Siga.

Belongs within: Tenebrionidae.

The Ulomini are a pantropical tribe of tenebrionid beetles found in well-rotted wood in forests. The Australian Typhluloma inops lacks eyes and may be a myrmecophile (Matthews & Bouchard 2008).

Characters (from Matthews & Bouchard 2008): Oblong, parallel-sided, convex, glabrous, piceous. Total length 4–20 mm. Edge of clypeus more or less straight in middle, clypeolabral membrane often exposed, at least in middle. Eyes (if present) reniform. Labrum trapezoidal, narrower basally, medial tormal arms of epipharynx directed somewhat anterad, expanded at their mesal extremities into very long anterior and shorter posterior extensions, epitorma absent, setal pattern symmetrical. Antennae short, reaching halfway or less to base of prothorax, segments gradually widening distally, transverse, distal edges of preapical segments bearing a circle of placoid sensoria, apical ones with stellate compound sensoria. Corpotentorium a little curved upwards. Mandibles apically bi- or tridentate, with short slender scissorial lobe and large mola, surface of mola with coarse teeth, not finely striate. Maxilla with two lacinial unci, one of which is laminar, palpi with apical segment obliquely securiform. Pronotum subquadrate, disc evenly convex or often anteriorly excavate in male. Prosternum moderately long before coxae. Procoxal cavities internally closed. Mesocoxal cavities closed by the ventrites. Metendosternite with laminae. Elytra with nine striae and scutellary striole, humeral angles dentate, epipleura not reaching apex, latter excised uncovering part of pygidium. Wings with venation unmodified, apical membrane up to one-third of wing length, without medial or apical flecks. Legs with pro- and mesotibiae fossorial, flattened and with dentate or spinulose outer edges, tarsomeres not lobed, plantar surfaces with moderately dense setae. Abdomen with last tergite partly exposed by excised elytral apices. Defensive gland reservoirs long, contracting to narrow bases, with large basomedial pouches, without common volume. Ovipositor shortened, with paraprocts shorter than coxites, gonostyles small and apical in position, paraproct baculi oblique, coxite baculi transverse. Female tract without bursa, with single coiled spermatheca attached to base of long slender accessory gland, without vaginal sclerites. Aedeagus with curved basal piece much longer than apical, its sides fused ventrally to form flattened tube, alae formed as extensions of apical piece.

<==Ulomini [Ulomina]
|--NeopsectropusMB08
|--Eutochia LeConte 1862 (see below for synonymy)AT02
|--Achthosus Pascoe 1863 [=Acthosus (l. c.)]MB08
| `--*A. westwoodi Pascoe 1863 [incl. A. insularis Lea 1908]MB08
|--Typhluloma Lea 1912MB08
| `--*T. inops Lea 1912MB08
|--Cenoscelis Wollaston 1867MB08
| `--‘Uloma’ pulla Erichson 1843 [incl. *Cenoscelis tibialis Wollaston 1867]MB08
`--Uloma Dejean 1821 [incl. Melasia Mulsant 1856, Prioscelida White 1846]AT02
| i. s.: U. cornutaR26
| U. janthina Dupont in Boisduval 1835B35
| U. longulaAT02
| U. nepalica Kaszab 1977K77
| U. perroudiiC01
| U. prehimalayica Kaszab 1975K77
| U. rubripes Hope 1831 [incl. U. orientalis Laporte de Castelnau 1840]K77
| U. westwoodiB70
|--U. (Uloma)MB08
| |--*U. (U.) culinaris (Linnaeus 1758)MB08 [=Tenebrio culinarisMB08, Diaperis (Phaleria) culinarisG20, L02]
| |--U. (U.) apteroides Kaszab 1982MB08
| |--U. (U.) australica Kaszab 1982MB08
| |--U. (U.) bituberosa Kirsch 1875 [incl. U. sydneyana Kaszab 1982]MB08
| |--U. (U.) blairi Kaszab 1982MB08
| |--U. (U.) brunnea (Carter 1906) [=Acthosus brunneus]MB08
| |--U. (U.) carteri Kaszab 1982MB08
| |--U. (U.) caviceps Gebien 1920MB08
| |--U. (U.) claudiei Kaszab 1982MB08
| |--U. (U.) clypeata Kaszab 1982MB08
| |--U. (U.) consentanea Perroud 1864 [incl. U. depressa Pascoe 1866]MB08
| |--U. (U.) impressicollisLC40
| |--U. (U.) kimberleyana Kaszab 1982MB08
| |--U. (U.) latior Carter 1926MB08
| |--U. (U.) lawrencei Kaszab 1982MB08
| |--U. (U.) leai Kaszab 1982MB08
| |--U. (U.) longipes Kaszab 1982MB08
| |--U. (U.) major Laporte de Castelnau 1840LC40
| |--U. (U.) marginatoides Kaszab 1982MB08
| |--U. (U.) neboissi Kaszab 1982MB08
| |--U. (U.) opacoides Kaszab 1982MB08
| |--U. (U.) orientalis Laporte de Castelnau 1840LC40
| |--U. (U.) orientaloides Kaszab 1982MB08
| |--U. (U.) pecki Kaszab 1982MB08
| |--U. (U.) planimentoides Kaszab 1982MB08
| |--U. (U.) queenslandica Kaszab 1982MB08
| |--U. (U.) retusa Fab. 1801LC40, FS90
| |--U. (U.) rubens Laporte de Castelnau 1840LC40
| |--U. (U.) sanguinipes (Fabricius 1775) [=Tenebrio sanguinipes; incl. Achthosus laticornis Pascoe 1869]MB08
| |--U. (U.) sexdecimlineata Montrouzier 1860 [incl. U. ovalis Perroud 1864]MB08
| |--U. (U.) tenebrionoides (White 1846) [=*Prioscelida tenebrionoides]MB08
| `--U. (U.) tibialis Kaszab 1982MB08
|--U. (Apterulomoides Kaszab 1982)MB08
| `--U. (*A.) rotundipennis Kaszab 1982MB08
|--U. (Ceratupis) nigerrimaLC40
`--U. (Uleda) diaperoides Laporte de Castelnau 1840LC40

Eutochia LeConte 1862 [incl. Aniara Lacordaire 1859, Aniarus Gemminger & Harold 1870, Delpygus LeConte 1866, Holaniara Fairmaire 1871]AT02

*Type species of generic name indicated

References

[AT02] Aalbu, R. L., C. A. Triplehorn, J. M. Campbell, K. W. Brown, R. E. Somerby & D. B. Thomas. 2002. Tenebrionidae Latreille 1802. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 463–509. CRC Press.

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d’Urville. Faune entomologique de l’océan Pacifique, avec l’illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[B70] Britton, E. B. 1970. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 495–621. Melbourne University Press.

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[FS90] Fleutiaux, E., & A. Sallé. 1890. Liste des coléoptères de la Guadeloupe et descriptions d’espèces nouvelles. Annales de la Société Entomologique de France, 6e série 9: 351–484.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[K77] Kaszab, Z. 1977. Tenebrionidae der Nepal-Expeditionen von Dr. J. Martens (1969–1974) (Insecta: Coleoptera). Senckenbergiana Biologica 57 (4–6): 241–283.

[LC40] Laporte de Castelnau, F. 1840. Histoire Naturelle des Insectes Coléoptères vol. 2. P. Duménil: Paris.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[MB08] Matthews, E. G., & P. Bouchard. 2008. Tenebrionid Beetles of Australia: Descriptions of tribes, keys to genera, catalogue of species. Australian Biological Resources Study: Canberra.

[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.

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