Isanda coronata, copyright Femorale.

Belongs within: Trochidae.

The Umboniinae are a group of small top shells found mostly on soft substrates in shallower waters. Genera related to Umbonium and Lirularia have a monopectinate ctenidium but the ctenidium remains bipectinate in basal genera such as Monilea and Ethalia (Herbert 1992).

Characters (from Herbert 1992): Radula with teeth of central field reduced, marginals well-developed and numerous, latero-marginal plate present; protoconch small (<250 µm in diameter) and beaked; teleoconch often with scratch-like axial microsculpture; snout with subterminal papillae; forehead frequently narrow or absent and snout reduced in size; eye stalks more or less elongate, curving up under shell margin in life, eyes usually large, terminal; left neck lobe variously modified to form particle exclusion filter, sometimes greatly hypertrophied; foot anteriorly bifid; ctenidium long, extending whole length of mantle cavity; ctenidial filaments elongate, with swollen tips and prominent bursicles, filaments on left side of ctenidial axis reduced or absent; afferent ctenidial membrane long, ctenidium sometimes attached to mantle skirt throughout its length.

Umboniinae [Bankiviini, Monileini, Rotelladae, Rotellinae, Umboniidae, Umboniini]BR17
| i. s.: ‘Minolia’ edithae Melvill 1891H92
| ‘Minolia’ biangulosa (Adams 1855)H92
| ‘Solariella’ dereimsi Dollfus 1911H92
| ‘Solariella’ canaliculata Smith 1872H92
|--Callumbonella Thiele 1924 [=Umbotrochus Thiele 1924 non Perner 1903]KC60
| `--*C. gorgonarum (Fischer 1883) [=Gibbula gorgonarum, *Umbotrochus gorgonarum]KC60
|--Inkaba Herbert 1992H92
| `--*I. tonga Herbert 1992H92
|--Zethalia Finlay 1926H92 [incl. Ethaliopsis Cossmann 1918 non Schepman 1908KC60]
| `--*Z. zelandicum (Adams 1854)P61 (see below for synonymy)
|--Bankivia Krauss 1848BR05
| `--*B. fasciata (Menke 1830)W93 (see below for synonymy)
|--Archiminolia Iredale 1929W93
| `--*A. oleacea (Hedley & Petterd 1907)W93 [=Monilea oleaceaKC60, Isanda (*Archiminolia) oleaceaKC60]
|--Parminolia Iredale 1929W93
| `--*P. apicina (Gould 1861)W93 (see below for synonymy)
|--Rossiteria Brazier 1895W93 [=Solanderia Fischer in Kiener 1879 non Duchassaing & Michelin 1846KC60]
| `--*R. nucleus (Philippi 1849)W93 [=Trochus nucleusW93, Monilea (*Rossiteria) nucleaKC60]
|--Antisolarium Finlay 1926H92
| |--*A. egenum (Gould 1849)P61 (see below for synonymy)
| `--A. stoliczkai (Zittel 1865) [=Solariella stoliczkai]F27
|--Vanitrochus Iredale 1929W93 [incl. Conotrochus Pilsbry 1889 non Schröter 1863KC60]
| |--*V. tragema (Melvill & Standen 1896)W93 [=Solariella tragemaW93, Isanda (*Vanitrochus) tragemaKC60]
| `--‘Trochus’ semiustus Fischer 1879W93
|--Leiopyrga Adams & Adams 1863W93
| |--*L. lineolaris (Gould 1861)W93, KC60 (see below for synonymy)
| |--L. cingulata (Adams 1863)W93 [=Cantharidus cingulatusH09]
| `--L. octona (Tate 1891) [incl. L. octona problematica (Iredale 1924)]W93
|--Pseudominolia Herbert 1992H92
| |--*P. splendens (Sowerby 1897) [=Solariella splendens; incl. Minolia variegata Odhner 1919]H92
| |--‘Solariella’ algoensis Thiele 1925 (see below for synonymy)H92
| `--‘Minolia’ climacota Melvill 1897H92
|--Talopena Iredale 1918H92
| |--*T. incerta (Iredale 1913)H92 [=Monilea incertaKC60]
| |--T. gloriola Iredale 1929H92
| |--T. lifuana (Fischer 1878)H92 [=Trochus lifuanusH92, Monilea lifuanaH09]
| |--T. vernicosa (Gould 1861)MG-H11
| `--T. vitiliginea (Menke 1843)F27, H09 [=Trochus vitiligineusH09, Monilea vitiligineaH09]
|--Isanda Adams & Adams 1854 [Isandini]BR05
| | i. s.: I. hornungi (Bisacchi 1931)H92
| |--*I. (Isanda) coronata Adams 1854KC60
| |--I. (Conominolia Finlay 1927)KC60
| | |--I. (*C.) conica (Marshall 1917) [=Heliacus conicus]KC60
| | `--‘Conominolia’ sulcatina (Suter 1917) [=Solariella sulcatina]F27
| |--I. (Umbonella Adams 1863)KC60
| | `--I. (*U.) murrea (Reeve 1848) [=Turbo murreus]KC60
| `--I. (Waisiolia Beets 1942)KC60
| `--I. (*W.) jucanda Beets 1942KC60
|--Umbonium Link 1807H92 [=Globulus Schumacher 1817KC60, Rotella Lamarck 1822BR05]
| | i. s.: U. costatum (Valenciennes in Kiener 1838–1839)H92
| | U. eloiseae Dance, Moolenbeek & Dekker 1992BC01
| | *Rotella’ lineolata Lamarck 1822BR17
| | ‘Rotella’ neozelanicaH86
| |--*U. (U.) vestiarium (Linnaeus 1758) [=Trochus vestiarius, *Globulus vestiarius, Rotella vestiaria]KC60
| |--U. (Ethaliella Pilsbry 1905)KC60
| | `--U. (*E.) floccata (Sowerby 1903) [=Ethalia floccata]KC60
| |--U. (Protorotella Makiyama 1925)KC60
| | `--U. (*P.) yuantaniensis (Makiyama 1925) [=*Protorotella yuantaniensis]KC60
| `--U. (Suchium Makiyama 1925)KC60
| `--U. (*S.) suchiense Yokoyama 1923KC60
|--Lirularia Dall 1909H92, BR05 [Lirulariinae]
| |--*L. lirulata (Carpenter 1863)BR17 [=Margarites lirulataBR17; incl. Margarita lirulata var. conicaC64]
| | |--L. l. lirulataO27
| | |--L. l. obsoleta (Carpenter 1864) [=Margarites lirulata obsoleta]O27
| | `--L. l. subelevata (Carpenter 1864) [=Margarites lirulata subelevata]O27
| |--L. acuticostata (Carpenter 1864)O27 [=Margarites (Lirularia) acuticostatusB56]
| |--L. althorpensis (Dall 1919) [=Margarites (Lirularia) althorpensis]O27
| |--L. antoniae Rubio & Rolán 1997BC01
| |--L. inflatula (Dall 1920) [=Margarites (Lirularia) inflatula]O27
| |--L. iridescens (Schrenck 1863)H84
| |--L. lacunata (Carpenter 1864) [=Gibbula lacunata, Margarites (Lirularia) lacunata]O27
| |--L. pedroana [=Margarites (Lirularia) pedroana]B56
| |--L. smithi (Bartsch 1927) [=Margarites (Lirularia) smithi]O27
| `--L. succincta (Carpenter 1864) [=Gibbula succincta, Margarites (Lirularia) succincta]O27
|--Ethminolia Iredale 1924 [incl. Conotalopia Iredale 1929, Sericominolia Kuroda & Habe 1952]H92
| |--*E. probabilis Iredale 1924H92
| |--E. durbanensis (Kilburn 1977) [=Solariella durbanensis]H92
| |--E. eilikrines (Melvill 1891)H92
| |--E. elveri Cotton & Godfrey 1938W93
| |--E. eudeli (Deshayes 1863) [=Trochus eudeli]H92
| |--E. gravieri (Lamy 1909) [=Cyclostrema gravieri]H92
| |--‘Minolia’ henniana Melvill 1891H92 [=Isanda (*Conotalopia) hennianaKC60, Monilea hennianaH09]
| |--‘Solariella’ illustris Sturany 1903H92
| |--E. impressa (Nevill & Nevill 1869) [=Solarium impressum, Minolia impressa; incl. M. eucoronata Sowerby 1905]H92
| |--E. nektonica (Okutani 1961) [=Solariella nektonica; incl. S. nektonica tajimensis Okutani & Sakurai 1968]H92
| |--E. sculpta (Sowerby 1897) [=Solariella sculpta, Minolia sculpta]H92
| |--E. stearnsii (Pilsbry 1895) [=Minolia stearnsii, *Sericominolia stearnsii, Solariella stearnsi]H92
| |--‘Conotalopia’ tropicalis (Hedley 1907)W93
| `--E. vitiliginea (Menke 1843) [incl. E. tasmanica Tenison Woods 1877]W93
|--Ethalia Adams & Adams 1854 [incl. Liotrochus Fischer 1878]H92
| |--*E. guamensis (Quoy & Gaimard 1834)H92 (see below for synonymy)
| | |--E. g. guamensisH92
| | `--E. g. selenomphala Pilsbry 1905H92
| |--E. bysma Herbert 1992H92
| |--E. carneolata Melvill 1897 [=Rotella carneolata; incl. E. carneolata var. rubrostrigata Melvill 1904]H92
| |--E. electra Herbert 1992H92
| |--E. gilchristae Herbert 1992H92
| |--E. lamprus (Watson 1880) [=Trochus (Solariella) lamprus]H92
| |--E. minolina Melvill 1897 [incl. E. minolina var. infralaevior Schepman 1908]H92
| |--E. minuta [=Vitrinella minuta]C64
| |--E. pulchella (Adams 1855) [=Isanda pulchella]H09
| |--E. regularis [=Vitrinella regularis]C64
| |--E. striolata (Adams 1855) [incl. E. trilobata Sowerby in Reeve 1875-1878]H92
| |--E. supravallata Carpenter 1864C64
| `--‘Vitrinella’ valvatoidesC64
`--Monilea Swainson 1840 [=Talopia Gray 1842; Talopiidae, Talopiini]BR05
| i. s.: M. angulata Adams 1851H08
| M. belcheri Philippi 1849W93
| M. corrugata [=Trochus corrugatus]B79
| M. glaphyrella (Melvill & Standen 1895) [=Minolia glaphyrella]H09
| M. lentiginosa Adams 1851W93
| M. morti Iredale 1929W93
| M. patricia (Philippi 1851)H92
| M. philippiana Dunker 1871WBS-S15
| M. philippii Adams 1855 [=Ethalia philippii]H92
| M. pudibunda (Fischer 1878) [=Trochus pudibundus]H09
| M. rosea Tenison Woods 1876TW76
| M. stoliczkaiH86
| M. tropicalis Hedley 1907H09
| M. vernicosa Gould 1861H92
|--*M. (Monilea) callifera (Lamarck 1822)KC60 [=Trochus calliferusKC60, *Talopia calliferaBR17]
`--M. (Priotrochus Fischer 1879)KC60, H92 [=Aphanotrochus Martens 1880KC60]
`--M. (*P.) obscura (Wood 1828) [=Trochus obscurus, *Aphanotrochus obscurus]KC60

*Antisolarium egenum (Gould 1849)P61 [=Solarium (Torinia) egenumH92, Minolia egenaF27, Solariella egenaF27; incl. Margarita dilecta Adams 1855H92, Solariella dilectaH92, Trochus (Solariella) dilectusH92]

*Bankivia (Bankivia) fasciata (Menke 1830)W93 [=Phasianella fasciataKC60, Cantharidus fasciatusF27; incl. Bankivia lugubrisH08, B. varians Krauss 1848KC60]

*Leiopyrga lineolaris (Gould 1861)W93, KC60 [=Cantharidus lineolarisKC60, Bankivia (Leiopyrga) lineolarisKC60; incl. Leiopyrga picturata Adams & Adams 1863KC60, Cantharidus picturatusF27]

*Ethalia guamensis (Quoy & Gaimard 1834)H92 [=Rotella guamensisH09, Umbonium (*Ethalia) guamenseKC60; incl. R. montrouzieri Souverbie 1860H92, *Liotrochus montrouzieriH92]

*Parminolia apicina (Gould 1861)W93 [=Monilea apicinaW93, Isanda (Parminolia) apicinaKC60; incl. Minolia agapeta Melvill & Standen 1869KC60, W93, Parminolia agapetaKC60]

‘Solariella’ algoensis Thiele 1925 [incl. Margarita articulata Gould 1861 non Adams 1853, Gibbula articulata, Minolia articulata, Pseudominolia articulata, Mi. bleeki Thiele 1925]H92

*Zethalia zelandica (Adams 1854)P61 [=Umbonium zelandicumP61, Ethalia zelandica Rousseau 1854F27; incl. Globulus anguliferus Philippi 1854 non Sowerby 1840F27, *Ethaliopsis anguliferaF27, Umbonium anguliferumF27]

*Type species of generic name indicated


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