Dinychus sp., copyright Scott Justis.

Belongs within: Uropodina.
Contains: Oplitis, Trachyuropoda, Nenteria, Uroobovella, Trichouropoda, Uropoda, Discourella, Trichocylliba.

The Uropodoidea are a diverse group of mites found in soil, humus, animal nests or decaying organic matter. Species may feed on fungi or organic debris or they may be predators on other small animals such as nematodes or insect larvae (Lindquist et al. 2009).

The classification of uropodoids has been somewhat chaotic in the past, largely due to debate on how much systematic weight should be allowed to characters that may not be expressed at all stages of the life cycle. In species of the family Uroactiniidae, the apex of the fixed cheliceral digit bears a large flower- or mushroom-shaped sensory organ and there are terminal teeth on the corniculi. A distinct internal sclerotised node is associated with the levator tendon of the chelicera in species of Trematuridae, Dinychidae, Trachyuropodidae and Oplitidae but not in Metagynella and Uropodidae. In Metagynella the hypostomatic setae h1 are short and spiniform, and the female genital shield is located behind coxae IV; Uropodidae usually have long and setiform setae h1 and the female genital shield is between coxae II–IV. Dinychidae, Trachyuropodidae and Oplitidae possess short marginal fimbriations and/or moustache-like distal excrescences on the internal malae that are absent in Trematuridae. In Dinychidae, the peritremes are well developed, often forming distinctive serpentine patterns, and the fixed cheliceral digit usually bears an apical ‘finger’ extending well past the movable digit. Trachyuropodidae have a strongly ornamented dorsal shield marked by pits and ridges, often with numerous T-shaped setae (Lindquist et al. 2009).

Characters (from Lindquist et al. 2009): Coxae I broad, covering or partially covering narrow tritosternal base, vertex and camerostome well developed, pedofossae often present; palpgenu with 4–5 setae. Dorsal shield of adults smooth or variously ornamented, sometimes with deep pits and ridges, entire or with a posterior pygidial element, free or contiguous with holoventral shield, fused anteriorly to peritrematic elements and often bordered by marginal scutella or coalesced marginal shields; pygidial shield, when present, may be narrow to broad, sometimes subdivided, with or without setae. Chelicerae elongate, sometimes appearing four-segmented, without lyrifissure, terminal digits usually small and often bearing elaborate sensory structures, pilus dentilis present or absent, levator tendon with or without a sclerotized node. Peritremes well developed and often convoluted or serpentine, tritosternal laciniae fused basally and often subdivided distally. Femur I with 8–9 setae, genua III–IV without (or occasionally with 1) ventral setae, tibia I with only 3 dorsal and 2 ventral setae (typically 1 1/1, 2/1 1). Female genital opening covered by a single nude epigynal shield, paraanal elements absent. Male with holoventral shield, genital opening between coxae II–IV (rarely postcoxal), covered by 1–2 valves; protonymph with separate or fused podonotal and mesonotal elements; phoretic deutonymphs may attach to host via strand produced by anal pedicel.

<==Uropodoidea [Prodinychini]LKW09
    |--Clausiadinychus Sellnick 1930KA90, FH93 [ClausiadinychidaeFH93]
    |    `--C. cristatus Sellnick 1930FH93
    |--Metagynella Berlese in Trouessart & Berlese 1919FH93 (see below for synonymy)
    |    |--*M. paradoxa Berlese 1919ET79
    |    |--M. lindquisti Hirschmann 1979FH93
    |    |--M. mexicana Hirschmann 1979FH93
    |    |--M. moseri Hirschmann 1975FH93
    |    `--M. parvula Camin 1953FH93
    |    |--Dentibaiulus Schuster 1962FH93
    |    |--Dicornutophorus Hirschmann 1979FH93
    |    |--Mayaphorus Hirschmann 1979FH93
    |    `--Eutrachytes Berlese 1914S61, FH93
    |         `--E. gaieri Schweizer 1961S61
    |    |--Allodinychus Tragardh 1943FH93
    |    |--Trachyxenura Leitner 1947FH93
    |    `--Prodinychus Berlese 1917KA90, FH93
    |         |--*P. fimicolus (Berlese 1903) [=Dinychus fimicolus]ET79
    |         |--P. americanus (Banks 1904) [=Dinychus americanus; incl. Uroobovella sellnickiamericana Hirschmann 1979]FH93
    |         |--P. flagellifer (Berlese 1910)FH93 [=Dinychus flagelliferFH93, Allodinychus flagelligerS61]
    |         |--P. hummelincki Sellnick 1963FH93
    |         `--P. tetraphyllus (Berlese 1903)FH93 [=Dinychus tetraphyllusFH93, Phyllodinychus tetraphyllusS61]
    |    |--OplitisLKW09
    |    |--Berleseoplitis Hirschmann 1979FH93
    |    |--Cariboplitis Sellnick 1963FH93
    |    |--Chelonuropoda Sellnick 1954FH93
    |    |--Marginura Sellnick 1926FH93
    |    |--Sellnickoplitis Hirschmann 1979FH93
    |    |--Stammeroplitis Hirschmann 1979FH93
    |    |--Uropectinia Kneissl 1908FH93
    |    |--Uroplitana Sellnick 1926FH93
    |    |--Woelkeoplitis Hirschmann 1979FH93
    |    `--Womersleyoplitis Hirschmann 1979FH93
    |--Uroactiniidae [Uroactiniinae]LKW09
    |    |--Chiropturopoda Sellnick 1958FH93
    |    |--Centrouropoda Berlese 1916FH93
    |    |    `--C. rackae Hirschmann 1975FH93
    |    `--Uroactinia Hirschmann & Zirngiebl-Nicol 1964H98
    |         |--*U. consanguinea [=Uropoda consanguinea]D81
    |         |--U. agitans (Banks in Newell 1908)LKW09, FH93 [=Uropoda agitansFH93]
    |         |--U. anchor (Trouessart 1902) [=Uropoda anchor; incl. Fuscuropoda hippocrepoides Vitzthum 1939]FH93
    |         |--U. centroamericana (Stoll 1893) [=Uropoda centroamericana]FH93
    |         |--U. domrowi Hirschmann 1990H98
    |         |--U. hippocrepea (Berlese 1918) [=Fuscuropoda hippocrepea]FH93
    |         |--U. hirschmanni Hiramatsu 1978FH93
    |         |--U. ovata (Fox 1948) [=Fuscuropoda ovata]FH93
    |         `--U. rarosaeVKC07
    |--Phymatodiscus Berlese 1917VKC07, FH93 [PhymatodiscidaeFH93]
    |--Trachyuropodidae [Trachyuropodina, Trachyuropodini, Trachyuropodoidea]LKW09
    |    |--TrachyuropodaLKW09
    |    |--Arculatatrachys Hirschmann 1979FH93
    |    |--Bostocktrachys Hirschmann 1979FH93
    |    |--Castritrachys Hirschmann 1979FH93
    |    |--Cephalojanetia Willmann 1951FH93
    |    |--Cephalouropoda Berlese 1903FH93
    |    |--Cristicepstrachys Hirschmann 1979FH93
    |    |--Excavatatrachys Hirschmann 1979FH93
    |    |--Foliatrachys Hirschmann 1979FH93
    |    |--Graecatrachys Hirschmann 1979FH93
    |    |--Lindquisttrachys Hirschmann 1979FH93
    |    |--Magnatrachys Hirschmann 1979FH93
    |    |--Origmatrachys Hirschmann 1979FH93
    |    |--Trogulotrachys Hirschmann 1979FH93
    |    |--Leonardiella Berlese 1904FH93
    |    |    `--*L. canestriniana (Berlese 1891)ET79, FH93 [=Uropoda canestrinianaFH93]
    |    `--Urojanetia Berlese 1913 [incl. Janetiella Berlese 1904 (preoc.)]FH93
    |         |--*U. coccinea (Michael 1891)E57 [=Uropoda coccineaE57, Trachyuropoda coccineaS22]
    |         `--U. cristipesS61
    |    |--NenteriaKA90
    |    |--Calurodiscus Radford 1950FH93
    |    |--Castrinenteria Hirschmann 1979FH93
    |    |--Stylinenteria Hirschmann 1979FH93
    |    |--Austrodinychus Trägårdh 1952FH93
    |    |    |--A. canadiensis (Hirschmann 1978) [=Nenteria canadiensis]FH93
    |    |    |--A. chiapasus (Hirschmann 1978) [=Nenteria chiapasa]FH93
    |    |    |--A. fungivorus (Hirschmann 1978) [=Nenteria fungivora]FH93
    |    |    |--A. howdeni (Hirschmann 1978) [=Nenteria howdeni]FH93
    |    |    |--A. martini (Hirschmann 1978) [=Nenteria martini]FH93
    |    |    |--A. mexicanus (Hirschmann 1978) [=Nenteria mexicana]FH93
    |    |    |--A. moseri (Hirschmann 1972) [=Nenteria moseri]FH93
    |    |    `--A. sellnicki (Hirschmann & Wiśniewski 1985) [=Nenteria sellnicki]FH93
    |    `--Ruhmnenteria Hirschmann 1979FH93
    |         |--R. bretonensis (Hirschmann 1978) [=Nenteria bretonensis]FH93
    |         |--R. chihuahuaensis (Hirschmann 1978) [=Nenteria chihuahuaensis]FH93
    |         |--R. durangoensis (Hirschmann 1978) [=Nenteria durangoensis]FH93
    |         |--R. eutamiasae (Hirschmann 1978) [=Nenteria eutamiasae]FH93
    |         |--R. microti (Hirschmann 1978) [=Nenteria microti]FH93
    |         |--R. orghidani (Huţu 1977) [=Nenteria orghidani]FH93
    |         |--R. potosi (Hirschmann 1978) [=Nenteria potosi]FH93
    |         `--R. saltoensis (Hirschmann 1978) [=Nenteria saltoensis]FH93
    |--Dinychidae [Dinychini, Macrodinychidae, Urodiaspididae]LKW09
    |    |--UroobovellaLKW09
    |    |--Diurodinychus Berlese 1916FH93
    |    |--Phyllodinychus Trägårdh 1943FH93
    |    |--Lindquistidiaspis Hirschmann 1984FH93
    |    |    `--L. lindquisti (Hirschmann 1979) [=Urodiaspis lindquisti]FH93
    |    |--Urodiaspis Berlese 1916LKW09, FH93
    |    |    |--*U. tecta (Kramer 1876)ET79 [=Notaspis tectusET79, Uropoda tectaE57]
    |    |    |--U. pannonica Willmann 1951K07
    |    |    `--U. walkeri Hirschmann & Zirngiebl-Nicol 1969FH93
    |    |--Crinitodiscus Sellnick 1931FH93
    |    |    `--C. (Oriendiscus)EU03
    |    |         |--C. (O.) pawlolowskiEU03
    |    |         `--C. (O.) rafalskiiEU03
    |    |--Dinychus Kramer 1886H98
    |    |    |--*D. perforatus Kramer 1882ET79
    |    |    |--D. greensladeae Bloszyk & Halliday 1995H98
    |    |    |--D. inermis (Koch 1841) [=Sejus inermis; incl. D. vartismithi Hull 1918]FH93
    |    |    |--D. micropunctatus Evans 1955FH93
    |    |    |--D. ornatus (Fox 1957) [=Phyllodynchus ornatus]FH93
    |    |    |--D. ovatus (Ewing 1909) [=Polyaspis ovatus]FH93
    |    |    |--D. ruseki Athias-Binche, Błoszyk & Olszanowski 1989FH93
    |    |    `--D. septentrionalis (Trägårdh 1943) [=Phyllodinychus septentrionalis]FH93
    |    |--Baloghmacrodinychus Hirschmann 1979FH93
    |    |--Bregetovamacrodinychus Hirschmann 1979FH93
    |    |--Loksamacrodinychus Hirschmann 1979FH93
    |    `--Macrodinychus Berlese 1917LKW09, H98
    |         |  i. s.: M. andrassyi Hirschmann 1975H98
    |         |         M. baloghi Hirschmann 1975H98
    |         |         M. parallelepipidusKGG07
    |         |--M. (Macrodinychus)FH93
    |         |    |--M. (M.) paraguayensisVKC07
    |         |    `--M. (M.) sellnicki Hirschmann & Zirngiebl-Nicol 1974FH93
    |         `--M. (Monomacrodinychus) multispinosus Sellnick 1973FH93
    |--Trematuridae [Trematurellidae, Trematurini]LKW09
    |    |--TrichouropodaLKW09
    |    |--Trematurella Trägårdh 1942FH93
    |    |--Dentidinychus Sellnick 1926FH93
    |    |--Gitodinychus Berlese 1917FH93
    |    |--Trematuroides Cooreman 1960FH93
    |    |--Trichodinychus Berlese 1917FH93
    |    |--Trichointerstructura Hirschmann 1979FH93
    |    |--Trichosociata Hirschmann 1979FH93
    |    |--Urolaelaps Berlese 1916FH93
    |    |--Urospina Sellnick 1931FH93
    |    |--Trematura Berlese 1917FH93
    |    |    |--*T. patavina (Canestrini 1885) [=Uropoda patavina]ET79
    |    |    `--T. jacksoniET79
    |    `--Oodinychus Berlese 1917FH93
    |         |--*O. janeti (Berlese 1904) [=Urodinychus janeti]ET79
    |         |--O. granulatus Sellnick 1963FH93
    |         |--O. jurassicus Schweizer 1961S61
    |         |--O. karawaiewi Berlese 1903S61
    |         |--O. margaritaensis Sellnick 1963FH93
    |         |--O. ovalis (Koch 1839)MS91 [=Uropoda ovalisS61]
    |         `--O. venezolanus Sellnick 1963FH93
    |--Urodinychidae [Urodinychini]FH93
    |    |--Appendiculobovella Hirschmann 1979FH93
    |    |--Calouropoda Berlese 1916FH93
    |    |--Dinychopsis Berlese 1916FH93
    |    |--Formosaobovella Hirschmann 1979FH93
    |    |--Indotrachytes Deb & Raychandhuri 1965FH93
    |    |--Olouropoda Berlese 1916FH93
    |    |--Paradinychus Berlese 1916FH93
    |    |    `--*P. venustus Berlese 1916B16
    |    |--Phaulotrachytes Oudemans 1913FH93
    |    |--Polyaspidiella Berlese 1910FH93
    |    |--Pulchellaobovella Hirschmann 1979FH93
    |    |--Styluropoda Trägåardh 1952FH93
    |    |--Urocyclella Berlese 1913FH93
    |    |--Urocylellopsis Willmann 1953FH93
    |    |--Dinychura Berlese 1913FH93
    |    |    |--*D. rectangula (Berl. 1913) [=Trachyuropoda (Urojanetia) rectangula]E57
    |    |    `--D. cordieri Berlese 1916 [=Discourella cordieri]ET79
    |    |--Vinicoloraobovella Hirschmann 1979FH93
    |    |    `--V. americana (Hirschmann 1972) (see below for synonymy)FH93
    |    |--Dendrouropoda Willmann 1959FH93
    |    |    `--D. valleiS79
    |    |--Neoseius Oudemans 1903FH93
    |    |    |--N. neptuni (Schuster 1958) [=Urosternella (Neoseius) neptuni]S58
    |    |    `--N. novusWP99
    |    |--Sellnickiobovella Hirschmann 1984FH93
    |    |    |--S. hilli (Sellnick 1970) [=Fuscuropoda hilli]FH93
    |    |    `--S. marmorea (Fox 1948) [=Fuscuropoda marmorea]FH93
    |    |--Urosternella Berlese 1903FH93
    |    |    `--U. cylindrica (Berlese 1913) [=Urodinychus cylindricus]FH93
    |    `--Urodinychus Berlese 1903ET79
    |         |--*U. carinatus (Berlese 1888) [=Uropoda carinata]ET79
    |         |--U. karawaiewiS22
    |         `--U. ovalisS22
    `--Uropodidae (see below for synonymy)LKW09
         |--Australiuropoda Hirschmann 1979FH93
         |--Ceyloniphorus Hirschmann 1979FH93
         |--Discotrachytes Berlese 1916FH93
         |--Foveolaturopoda Hirschmann 1979FH93
         |--Gibbauropoda Hirschmann 1979FH93
         |--Heterodinychus Berlese 1912FH93
         |--Interrupturopoda Hirschmann 1979FH93
         |--Jerzywisniewskia Hirschmann 1979FH93
         |--Karguropoda Hirschmann 1979FH93
         |--Laqueaturopoda Hirschmann 1979FH93
         |--Manuleatophorus Hirschmann 1979FH93
         |--Neobirophorus Hirschmann 1979FH93
         |--Multiporuropoda Hirschmann 1979FH93
         |--Odonturopoda Marais 1977FH93
         |--Penicillaturopoda Hirschmann 1979FH93
         |--Spiculaturopoda Hirschmann 1979FH93
         |--Spinosissuropoda Hirschmann 1979FH93
         |--Dentatadinychus Hirschmann 1979FH93
         |--Marginalidinychus Hirschmann 1979FH93
         |--Toplaidinychus Hirschmann 1979FH93
         |--Uropolyaspis Berlese 1903FH93
         |    `--*U. hamulifera (Michael 1894) [=Uropoda hamulifera]ET79
         |--Cyclacarus Ewing 1933FH93
         |    `--C. aberrans Ewing 1933FH93
         |--Kaszabjbaloghia Hirschmann 1973 [Kaszabjbaloghiidae]H98
         |    `--K. hirschmanni Hiramatsu 1978H98
         |--Olodiscus Berlese 1917FH93
         |    |--*O. integer (Berlese 1910) [=Discopoma integra]ET79
         |    `--O. minima Kramer 1882E57
         |--Metadinychus Berlese 1916FH93
         |    |--M. argasiformis Berlese 1916FH93
         |    |--M. exilis (Hiramatsu & Hirschmann 1979) [=Uropoda (Metadinychus) exilis]FH93
         |    `--M. pura (Hiramatsu & Hirschmann 1979) [=Uropoda (Metadinychus) pura]FH93
         |--Tuberdinychus Schweizer 1961FH93
         |    |--*T. subterranus (Schweizer 1922)S61 [=Urodinychus subterranusS22]
         |    |--T. fumicolus Schweizer 1961S61
         |    `--T. parvus Schweizer 1961S61
         |--Castriidinychus Hirschmann 1973H98
         |    |--C. baloghi Hirschmann 1975H98
         |    |--C. kaszabi Hirschmann 1975H98
         |    `--C. mahunkai Hirschmann 1975H98
         |--Diphaulocylliba Vitzthum 1925FH93
         |    |--*D. amplior (Berlese 1923) [=Phaulocylliba amplior]V25
         |    |--D. debilior Vitzthum 1925V25
         |    |--D. ingens Vitzthum 1925V25
         |    `--D. permagna (Canestrini 1897) [=Discopoma permagna]V25
         |--Phaulocylliba Berlese 1903FH93
         |    |--*P. ventricosa Berlese 1903V25
         |    |--P. inflata Berlese 1920V25
         |    |--P. littoralis (Trouessart 1902) [=Discopoma littorale; incl. P. berlesi Halbert 1915]V25
         |    |--P. misella (Berlese 1916) [=Cyllibano (Microcylliba) misella]V25
         |    |--P. orbicularis (Müller 1882)S61 [incl. Discopoma romana Canestrini & Canestrini 1882V25, P. romanaS61]
         |    `--P. virgata (Hull 1918) [=Cillibano virgata]V25
         |--Deraiophorus Canestrini 1897LKW09, H98
         |    |--D. australis Hirschmann 1973H98
         |    |--D. maya (Krantz 1969) [=Eutrachytes maya]FH93
         |    |--D. neobiroi Hirschmann 1973H98
         |    |--D. novaehollandiae (Domrow 1957) [=Urodiaspis novaehollandiae]H98
         |    |--D. simplicior (Domrow 1957) [=Eutrachytes simplicior]H98
         |    `--D. truncatus (Berlese 1888) [=Celaeno truncata]FH93
         |--Piracarus crozetensisP94
         |--Baloghurella Hirschmann 1979FH93
         |--Capitodiscus Vitzthum 1931FH93
         |--Clivosurella Hirschmann 1979FH93
         |--Comydinychus Berlese 1917FH93
         |--Cosmogynurella Hirschmann 1979FH93
         |--Formosaurella Hirschmann 1979FH93
         |--Gerlachurella Hirschmann 1979FH93
         |--Longicarinaturella Hirschmann 1979FH93
         |--Reticulaturella Hirschmann 1979FH93
         |--Rotundurella Hirschmann 1979FH93
         |--Stammerurella Hirschmann 1979FH93
         |--Antennequesoma Sellnick 1926FH93
         `--Hirtitrichocyllibula Hirschmann 1979FH93

Metagynella Berlese in Trouessart & Berlese 1919FH93 [incl. Metagynura Balogh 1943FH93; Metagynellidae, MetagynuridaeLKW09]

Uropodidae [Castriidinychidae, Cillibidae, Circocyllibidae, Coxequesomidae, Deraiophoridae, Discourellidae, Phaulodinychidae, Phaulodinychini, Planodiscidae, Thrichocyllibidae, Trichocyllibidae, Uropoden, Uropodinae, Uropodini]LKW09

Vinicoloraobovella americana (Hirschmann 1972) [=Uroobovella americana, Fuscuropoda ameriana Moser & Roton 1971 (n. n.); incl. U. neoamericana Hirschmann & Zirngiebl-Nicol 1972]FH93

*Type species of generic name indicated


[B16] Berlese, A. 1916. Centuria terza di Acari nuovi. Redia 12: 289–338.

[D81] Domrow, R. 1981. A small lizard stifled by phoretic deutonymphal mites (Uropodina). Acarologia 22 (3): 247–252.

[EU03] Ekiz, A. N., & R. Urhan. 2003. A new record of Uroobovella Berlese, 1903 (Acari: Uropodina) from Turkey with some ecological notes. Zoology in the Middle East 29: 105–111.

[E57] Evans, G. O. 1957. An introduction to the British Mesostigmata (Acarina) with keys to families and genera. Journal of the Linnean Society of London, Zoology 43: 203–259.

[ET79] Evans, G. O., & W. M. Till. 1979. Mesostigmatic mites of Britain and Ireland (Chelicerata: Acari-Parasitiformes). An introduction to their external morphology and classification. Transactions of the Zoological Society of London 35: 139–270.

[FH93] Farrier, M. H., & M. K. Hennessey. 1993. Soil-inhabiting and free-living Mesostigmata (Acari-Parasitiformes) from North America: an annotated checklist with bibliography and index. North Carolina Agricultural Research Service, North Carolina State University, Technical Bulletin 302: i–xvi, 1–408.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[K07] Kontschán, J. 2007. Some new records of Uropodina mites (Acari: Mesostigmata) from Croatia, Serbia and Montenegro with descriptions of two new species. Annales Historico-Naturales Musei Nationalis Hungarici 99: 177–188.

[KA90] Krantz, G. W., & B. D. Ainscough. 1990. Acarina: Mesostigmata (Gamasida). In: Dindal, D. L. (ed.) Soil Biology Guide pp. 583–665. John Wiley & Sones: New York.

[KGG07] Krantz, G. W., L. A. Gómez & V. E. Gonzalez. 2007. Parasitism in the Uropodina: a case history from Colombia. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 29–38. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[LKW09] Lindquist, E. E., G. W. Krantz & D. E. Walter. 2009. Order Mesostigmata. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 124–232. Texas Tech University Press.

[MS91] Miko, L., & M. Stanko. 1991. Small mammals as carriers of non-parasitic mites (Oribatida, Uropodina). In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 395–402. SPB Academic Publishing: The Hague.

[P94] Pugh, P. J. A. 1994. Non-indigenous Acari of Antarctica and the sub-Antarctic islands. Zoological Journal of the Linnean Society 110: 207–217.

[S58] Schuster, R. 1958. Neue terrestrische milben aus dem Mediterranen littoral. Vie et Milieu 9 (1): 88–109.

[S79] Schuster, R. 1979. Soil mites in the marine environment. Recent Advances in Acarology 1: 593–602.

[S22] Schweizer, J. 1922. Beitrag zur Kenntnis der terrestrischen Milbenfauna der Schweiz. Verhandl. Naturf. Ges. Basel 33: 23–112, 4 pls.

[S61] Schweizer, J. 1961. Die Landmilben der Schweiz (Mittelland, Jura und Alpen): Parasitiformes Reuter, mit 246 Arten und Varietäten und 268 meist kombinierten Originalzeichnungen. Denkschriften der Schweizerischen Naturforschenden Gesellschaft [Mémoires de la Société Helvétique des Sciences Naturelles] 84: i–vii, 1–207.

[VKC07] Vázquez, M. M., H. Klompen & C. I. Chargoy. 2007. Study of the Uropodina (Acari: Mesostigmata) in tropical forests of the Yucatan Peninsula, Mexico and Belize. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 43–50. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[V25] Vitzthum, H. 1925. Fauna sumatrensis. (Beitrag no. 5). Acarinae. Supplementa Entomologica 11: 1–79.

[WP99] Walter, D. E., & H. C. Proctor. 1999. Mites: Ecology, Evolution and Behaviour. CABI Publishing: Wallingford (UK).

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