Vanikoroidea

Balcis bulbula, from New Zealand Mollusca.

Belongs within: Littorinina.
Contains: Vanikoro, Eulima, AclisStilifer, Melanella, Curveulima, Mucronalia, Balcis (Balcis).

The Vanikoroidea are a group of carnivorous marine gastropods associated by molecular data. Members include the Eulimidae which live as parasites on echinoderms from which they feed by means of a piercing proboscis. Shells of eulimids are commonly translucent, allowing the internal viscera to be observed in life. The Vanikoridae have often globose shells with extremely large apertures and may be sessile at maturity. Gigantocapulus are large Cretaceous limpet-like shells of uncertain affinities found in association with bivalves.

Vanikoroidea [Eulimacea, Eulimoidea]BR17
|--Gigantocapulus Hayami & Kanie 1980 [Gigantocapulidae]BR17
| `--*G. giganteus (Schmidt 1873) [=Helcion giganteus]BR17
|--Vanikoridae [Merriidae, Naricidae, Naricinae, Vanicoroidae]BR17
| |--KorovaniaF27a
| |--VanikoroBR05
| `--Vanikoropsis Meek 1876SH12
| |--V. arktowskiana (Wilckens 1910)SH12
| |--V. decussata (Deshayes 1842)TTE93
| |--V. jackiBRW98
| `--V. stuartiF71
`--Eulimidae (see below for synonymy)BR17
|--Annulobalcis aurisflamma Simone & Martins 1995S11
|--MonogamusPC08
|--CostaclisBR05
|--EulimaBR05
|--Semistylifer pellucidus (Deshayes 1824)TTE93
|--AclisBR17
|--Ophiolamia armigeri Warén & Carney 1981C94
|--StiliferBR05
|--MelanellaBR05
|--CurveulimaW93
|--MucronaliaW93
|--Asterophila Randall & Heath 1912 [Asterophilidae]BR05
| `--*A. japonica Randall & Heath 1912BR17
|--Paedophoropus Ivanov 1933 [Paedophoropodidae]BR05
| `--*P. dicoelobius Ivanov 1933BR17
|--Entoconcha Müller 1852 [Cochlosolenia, Entoconchidae, Entoconchini]BR05
| `--*E. mirabilis Müller 1852BR17 [incl. Helicosyrinx parasita Baur 1864BK77]
|--Enteroxenos Bonnevie 1902 [Enteroxenidae, Enteroxeninae, Enteroxenini]BR05
| `--*E. oestergreni Bonnevie 1902BR17 [incl. Comenteroxenos parastichopi Tikasingh 1961BK77]
|--Margineulima Cossmann 1888DK08
| `--M. danae (Tenison Woods 1879)DK08
|--‘Pherusa’ Jeffreys 1869 nec Oken 1807 nec Leach 1814 [Pherusidae]BR05
| `--*P. gulsonae (Clark 1850) [=Chemnitzia gulsonae]BR17
|--Asterolamia Warén 1980W93
| `--*A. hians Warén 1980W93
|--Megadenus Rosén 1910W93
| `--*M. holothuricola Rosén 1910W93
|--Microstilifer Warén 1980W93
| `--*M. auricula (Hedley 1907) [=Stylifer auricula]W93
|--Teretianax Iredale 1918P61
| |--*T. suteri [=Scalenostoma suteri]P61
| `--T. pagoda Powell 1926P61
|--Entocolax Voigt 1888 [Cochlosyringia, Entocolacidae]BR05
| |--*E. ludwigii Voigt 1888BR17
| `--E. schwanwitschiBK77
|--Chileutomia Tate & Cossmann 1898P61
| |--*C. subvaricosa Tate 1898F27b
| `--C. neozelanica Powell 1940P61
|--Eulitoma Laseron 1955W93
| |--*E. nitens Laseron 1955W93
| `--E. castanea Laseron 1955W93
|--Hemiliostraca Pilsbry 1918W93
| |--H. peasei (Tryon 1886)W93 (see below for synonymy)
| `--H. sloaniW93
|--Leutzenia Warén 1980W93
| |--*L. asthenosomae Warén 1980W93
| `--L. toki Habe 1974W93
|--Peasistilifer Warén 1980W93
| |--*P. nitidula (Pease 1860)W93 (see below for synonymy)
| `--‘Chryseulima’ solitaria Laseron 1950W93
|--Pictobalcis Laseron 1955W93
| `--*P. articulata (Sowerby 1834) [=Eulima articulata]W93
|--Pulicicochlea Ponder & Gooding 1978W93
| |--*P. calamaris Ponder & Gooding 1978W93
| `--P. fusca Ponder & Gooding 1978W93
|--Sticteulima Laseron 1955 [incl. Lentigobalcis Habe 1961]W93
| |--*S. cameroni Laseron 1955W93
| `--*Lentigobalcis’ lentignosaW93
|--Trochostilifer Warén 1980W93
| |--*T. domus Warén 1980W93
| `--T. striatus (Hedley 1905) [incl. T. prionocidaricola Habe 1974]W93
|--Vitreobalcis Warén 1980W93
| |--*V. holdsworthi (Adams 1874) [=Apicalia holdsworthi]W93
| `--V. laevis Warén 1980W93
|--Echineulima Lutzen & Nielsen 1975W93
| |--*E. mittrei (Petit 1851) [=Stylifer mittrei]W93
| |--E. ponderi Warén 1980W93
| `--E. thanuumi Pilsbry 1921W93
|--Thyca Adams & Adams 1854 (see below for synonymy)W93
| |--*T. astericola (Adams & Reeve 1850) [=Pileopsis astericola]BR17
| |--T. crystallina Gould 1846 [incl. T. pellucida Kukenthal 1897]W93
| `--T. stellasteris Köhler & Vaney 1912W93
|--Pyramidelloides Nevill 1884W93
| |--*P. miranda (Adams 1861) (see below for synonymy)W93
| |--P. glaber Faber 1990BC01
| |--P. multicostatus Faber 1990BC01
| `--P. viticula Laseron 1956JG19
|--Cythna Carpenter 1864O27a
| |--*C. tumens [=Stylifer tumens]O27a
| |--C. albida Carpenter 1864O27a
| `--C. asteriaphilaC64
|--Haliella Monterosato 1873O27a
| |--*H. stenostomaO27a
| |--H. abyssicola Bartsch 1908O27a
| `--H. lomana Dall 1908O27a
|--Fusceulima Laseron 1955W93
| |--*F. jacksonensis Laseron 1955 [incl. F. flava Laseron 1955, F. sucina Laseron 1955]W93
| |--F. brunnea Tate 1887W93
| `--F. perexigua Tate & May 1900W93
|--Paramegadenus Humphreys & Lutzen 1972W93
| |--*P. arrhynchus [=Megadenus arrhynchus]W93
| |--P. incerta Warén 1980W93
| `--P. scutellicola Warén 1980W93
|--Sabinella Monterosato 1890 [incl. Eulimoda Laseron 1955]W93
| |--S. pachya [incl. Eulima piriformis, *S. piriformis]W93
| |--S. bakeri (Bartsch 1917) [=Melanella (Sabinella) bakeri]O27a
| |--S. munita (Hedley 1903) [=*Eulimoda munita]W93
| `--S. ptilocrinicola (Bartsch 1907) [=Melanella (Sabinella) ptilocrinicola]O27a
|--Scalenostoma Deshayes 1863 [incl. Hyperlia Pilsbry 1918, Stilimella Laseron 1955]W93
| |--*S. carinata Deshayes 1863W93
| |--*Hyperlia’ apiculataW93
| |--S. lodderae Petterd 1884 (see below for synonymy)W93
| `--S. striatum Hedley 1900H09
|--Strombiformis da Costa 1778 (nom. rej.) [Strombiformidae]BR05
| |--*S. glaber da Costa 1778BR17
| |--S. alaskensis Bartsch 1917O27a
| |--S. almo Bartsch 1917O27a
| |--S. auricincta Abbott 1958B59
| `--S. californicus Bartsch 1917O27a
|--Apicalia Adams 1862 [incl. Chryseulima Laseron 1955]W93
| |--*A. gibba Adams 1862W93
| |--A. angulata Warén 1981W93
| |--A. brazieri (Angas 1877) [=*Chryseulima brazieri; incl. A. immaculata Pritchard & Gatliff 1900]W93
| |--A. cicatricosa Warén 1981W93
| `--A. echinasteri Warén 1981W93
|--Stilapex Iredale 1925 [incl. Lambertia Souverbie 1869 (preoc.)]W93
| |--*S. lactarius Iredale 1925W93
| |--S. montrouzieri (Souverbie 1869) [=*Lambertia montrouzieri]W93
| |--S. ophiuraphila Habe 1976W93
| |--S. parva Schepman 1909W93
| `--S. thielei Sturany 1903W93
|--Parvioris Warén 1981W93
| |--*P. fulvescens (Adams 1866) [=Eulima fulvescens]W93
| |--P. australiensis Warén 1981W93
| |--P. equestris Koehler & Vaney 1912W93
| |--P. noumeae Warén 1981W93
| |--P. styliferoides Melvill & Standen 1901W93
| `--P. subobtusa (Laseron 1955) [=Curveulima subobtusa; incl. C. lata Laseron 1955]W93
|--Hypermastus Pilsbry 1899W93
| |--*H. coxi Pilsbry 1899W93
| |--H. colmani Warén & Crossland 1991W93
| |--H. georgiregis Cotton & Godfrey 1932W93
| |--H. mucronata (Sowerby 1866)W93
| |--H. placentae Warén & Crossland 1991W93
| `--H. williamsi Cotton & Godfrey 1932W93
|--Hebeulima Laseron 1955W93
| |--*H. inusta (Hedley 1906) [=Leiostraca inusta]W93
| |--H. columnaria May 1915W93
| |--H. crassiceps Laseron 1955W93
| |--H. fricata Hedley 1907W93
| |--H. kilcundae Gatliff & Gabriel 1914W93
| |--H. perexigua Tate & May 1900W93
| `--H. tumere Laseron 1955W93
|--Pelseneeria Koehler & Vaney 1908BR05 (see below for synonymy)
| |--*P. profunda Koehler & Vaney 1908BR17
| |--‘Hypermastus’ bountyensis Powell 1933P61 [=*Venustilifer bountyensisW93]
| |--P. brunneus (Tate 1888)W93
| |--P. mediaBK77
| |--‘Venustilifer’ secundus Powell 1940P61
| |--*Parastilifer’ sibogaeW93
| |--P. stimpsoni (Verrill 1882) [=Stilifer stimpsoni]BK77
| `--P. stylifera (Turton 1825)BK77 (see below for synonymy)
|--Niso Risso 1826O27a
| |--N. baueri Emerson 1965BC01
| |--N. brunneaF66
| |--N. emersoni McLean 1970BC01
| |--N. hipolitensis Bartsch 1917O27a
| |--N. lineata (Conrad 1841)P12
| |--N. lomana Bartsch 1917O27a
| |--N. psilo Tenison-Woods 1879F71
| `--N. splendidula (Sowerby 1834)PP78
|--Balcis Leach 1847P61
| | i. s.: B. columbianaB56
| | B. devians [=Eulima devians; incl. E. philippii Watson 1886]BK77
| | B. eburneaGO06
| | B. grippiB56
| | B. intermedia (Cantraine 1835)S11
| |--B. (Balcis)P61
| `--B. (Vitreolina Monterosato 1884)B56, D56
| |--‘Melania’ distorta [incl. Balcis arcuata, Eulima incurva, *Vitreolina incurva]D56
| |--‘Vitreolina’ curva (Monterosato 1974)O96
| |--‘Vitreolina’ levantina Oliverio, Buzzurro & Villa 1994O96
| |--B. (V.) thersites (Carpenter 1864)B56, O27a (see below for synonymy)
| `--B. (V.) titubans Berry 1956B56
|--Cyclonidea carina Laseron 1956JG19
`--Palisadia subulata Laseron 1956JG19

Balcis (Vitreolina) thersites (Carpenter 1864)B56, O27 [=Melanella (Balcis) thersitesO27; incl. Eulima bistortaO27, E. loweiO27]

Eulimidae [Aclidae, Aclididae, Aclidinae, Aclidoidea, Aclisidae, Aglossa, Entoconchida, Entoconchilla, Euliminae, Exoconchilla, Melanellacea, Melanellidae, Saccopallia, Stiliferidae, Stiliferoidei, Styliferida, Styliferidae]BR17

Hemiliostraca peasei (Tryon 1886)W93 [incl. Leiostraca distorta Pease 1861 (preoc.)W93, Eulima distortaJ79, *Hemiliostraca distortaW93, Subularia distortaK65, Eulima distorta var. gracilisJ79]

*Peasistilifer nitidula (Pease 1860)W93 [=Mucronalia nitidulaW93, Eulima nitidulaK65; incl. Peasistilifer gracilis Pease 1867W93, P. holothuricola Adam 1934W93, P. rosea Pease 1860W93]

Pelseneeria Koehler & Vaney 1908BR05 [incl. Parastilifer Ivanov 1952W93, Rosenia Nierstrasz 1913 non Waagen & Wentzel 1886BR05, Stylina Fleming 1828 non Lamarck 1816BR05, Turtonia Rosén 1910 non Alder 1848BR05, Venustilifer Powell 1939W93; Pelseneeridae, Pelseneeriidae, Roseniidae, Turtoniidae]

Pelseneeria stylifera (Turton 1825)BK77 [=Phasianella styliferaBK77, *Rosenia styliferaBR17, Stilifer styliferBK77, *Stylina styliferaBR17, *Turtonia styliferaBR17]

*Pyramidelloides miranda (Adams 1861) [=Rissoina miranda; incl. P. cylindrica Laseron 1956, P. pacifica Laseron 1956, P. turris Laseron 1956]W93

Scalenostoma lodderae Petterd 1884 [=*Stilimella lodderae; incl. Sc. crotaphis Watson 1886, Sc. petterdi Tate & May 1901, Sc. robusta Petterd 1884]W93

Thyca Adams & Adams 1854 [incl. Bessomia Berry 1959, Granulithyca Habe 1976, Kiramodulus Kuroda 1949; Thycidae, Thycinae]W93

*Type species of generic name indicated

References

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