Vertigo pusilla, copyright O. Gargominy.

Belongs within: Pupilloidea.

The Vertigininae are a group of snails with reduced tentacles and a penis lacking an appendix and with a simple retractor.

<==Vertigininae [Vertigineae, Vertigini]
|--Bothriopupa Pilsbry 1898BP90
| `--B. variolosaBP90
|--Sterkia Pilsbry 1898BP90
| `--S. eyrieseBP90
| |--S. e. eyrieseBP90
| `--S. e. rhoadsiBP90
`--Vertigo Müller 1773BR17
| i. s.: V. californicaBP90
| V. coloradensisPF15 [=V. gouldi coloradensisB65]
| |--V. c. coloradensisPF15
| `--V. c. arizonensisPF15
| V. diversidensE99
| V. gouldiiW79
| V. insertaWK13
| V. interferens (Deshayes 1863)TTE93
| V. meramecensis VanDevender 1979NC01
| V. miliumBP90
| V. nouletiE99
| V. ovataPF15
| V. oviformis Michaud in Deshayes 1863TTE93
| V. parvulaBP90
| V. simplexW79
| V. tridentataG59
| V. ventricosaG59
|--V. (Vertigo)G40
| |--*V. (V.) pusilla Müller 1774BR17 [incl. Pupa vertigoG40]
| `--V. (V.) angustior [incl. Turbo vertigo]G40
`--V. (Isthmia)G40
|--V. (I.) alpestrisG40
|--V. (I.) cylindrica [=Alaea cylindrica, Helix (Isthima) cylindrica; incl. Pupa minuta, P. minutissima]G40
|--V. (I.) edentula (see below for synonymy)G40
|--V. (I.) palustrisG40 (see below for synonymy)
|--V. (I.) pygmaea [=Pupa pygmaea; incl. V. vulgaris, Alaea vulgaris]G40
`--V. (I.) substriata (see below for synonymy)G40

Vertigo (Isthmia) edentula [=Pupa edentula, Jaminia edentula, Turbo edentulus; incl. Alaea nitida, Vertigo nitida, Turbo offtonensis, Alaea revoluta]G40

Vertigo (Isthmia) palustrisG40 [=Alaea palustrisG40; incl. Pupa antivertigoG40, Vertigo antivertigoH79, V. montagua msG40, Helix septemdentataG40]

Vertigo (Isthmia) substriata [=Alaea substriata, Pupa substriata; incl. Turbo sexdentatus, P. sexdentata, Vertigo sexdentata, V. similis]G40

*Type species of generic name indicated


[B65] Beetle, D. E. 1965. Molluscan fauna of some small ponds in Grand Teton National Park. Nautilus 78 (4): 125–130.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BP90] Burch, J. B., & T. A. Pearce. 1990. Terrestrial Gastropoda. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 201–309. John Wiley & Sones: New York.

[E99] Esu, D. 1999. Contribution to the knowledge of Neogene climate changes in western and central Europe by means of non-marine molluscs. In: J. Agustí, L. Rook & P. Andrews (eds) The Evolution of Neogene Terrestrial Ecosystems in Europe pp. 328–354. Cambridge University Press.

[G40] Gray, J. E. 1840. A Manual of the Land and Fresh-water Shells of the British Islands, with figures of each of the kinds. By William Turton, M.D. A new edition, thoroughly revised and much enlarged. Longman, Orme, Brown, Green, and Longmans: London.

[G59] Grimm, W. 1959. Land snails of Carroll County, Maryland. Nautilus 72 (4): 122–127.

[H79] Hey, W. C. 1879. Contributions to a better knowledge of the land and freshwater shells of Yorkshire. Journal of Conchology 2: 310–314.

[NC01] Nekola, J. C., & B. F. Coles. 2001. Systematics and ecology of Gastrocopta (Gastrocopta) rogersensis (Gastropoda: Pupillidae), a new species of land snail from the Midwest of the United States of America. Nautilus 115 (3): 105–114.

[PF15] Pilsbry, H. A., & J. H. Ferriss. 1915. Mollusca of the southwestern states, VII: the Dragoon, Mule, Santa Rita, Baboquivari, and Tucson Ranges, Arizona. Proceedings of the Academy of Natural Sciences of Philadelphia 67 (2): 363–418, pls 8–15.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

[W79] Walker, B. 1879. Catalogue of the shell-bearing Mollusca of Michigan. Journal of Conchology 2: 325–337.

[WK13] Whisson, C. S., & F. Köhler. 2013. Gastrocopta (Mollusca, Gastropoda, Pupillidae) in the Pilbara region of Western Australia. ZooKeys 261: 15–39.

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