Vombatomorphia

 Southern hairy-nosed wombat Lasiorhinus latifrons, from Blanchetown Allsorts.

Belongs within: Diprotodontia.
Contains: Diprotodontidae.

The Vombatomorphia include the living wombats (Vombatidae) and their fossil relatives. These include such diverse forms as the marsupial lions (Thylacoleonidae), the rhinoceros-like Diprotodontidae, and the large clawed herbivores of the Palorchestidae.

Vombatomorphia
    |--Thylacoleonidae [Wakaleoninae]LA02
    |    |--Priscileo Rauscher 1987LA02
    |    |    |--*P. pitikantensis Rauscher 1987LA02
    |    |    `--P. roskellyae Gillespie 1997LA02
    |    `--+--Wakaleo Clemens & Plane 1974LA02
    |       |    |--W. oldfieldi Clemens & Plane 1974AD82, LA02
    |       |    `--+--*W. vanderleueri Clemens & Plane 1974LA02, AD82, LA02
    |       |       `--W. alcootaensis Archer & Rich 1982AD82, LA02
    |       `--Thylacoleo Gervais 1852LA02 (see below for synonymy)
    |            |--T. hilli Pledge 1977AD82, LA02
    |            `--+--*T. carnifex Owen 1858LA02, AD82, LA02 (see below for synonymy)
    |               `--T. crassidentatus Bartholomai 1962AD82, LA02
    `--+--IlariidaeLA02
       |    |--Kuterintja Pledge 1987LA02
       |    |    `--*K. ngama Pledge 1987LA02
       |    `--Ilaria Tedford & Woodburne 1987LA02
       |         |--*I. illumidens Tedford & Woodburne 1987LA02
       |         `--I. lawsoni Tedford & Woodburne 1987LA02
       |--WynyardiidaeBA12
       |    |--Ayekaye jaredi Megirian et al. 2004BA12
       |    |--Wynyardia Spencer 1901LA02
       |    |    `--*W. bassiana Spencer 1901LA02
       |    |--Muramura Pledge 1987LA02
       |    |    |--*M. williamsi Pledge 1987LA02
       |    |    `--M. pinpensis Pledge 2003BA12
       |    `--Namilamadeta Rich & Archer 1979LA02
       |         |--*N. snideri Rich & Archer 1979LA02
       |         |--N. albivenator Pledge 2005BA12
       |         |--N. crassirostrum Pledge 2005BA12
       |         `--N. superior Pledge 2005BA12
       `--+--Marada Black 2007B07 [MaradidaeBA12]
          |    `--*M. arcanum Black 2007BA12
          `--+--DiprotodontoideaLA02
             |    |--DiprotodontidaeBA12
             |    `--PalorchestidaeLA02
             |         |--Propalorchestes Murray 1986LA02
             |         |    |--*P. novaculacephalus Murray 1986LA02
             |         |    `--P. ponticulus Murray 1990LA02
             |         `--Palorchestes Owen 1873LA02
             |              |--*P. azael Owen 1873LA02
             |              |--P. anulus Black 1997LA02
             |              |--P. painei Woodburne 1967LA02
             |              |--P. parvus De Vis 1895LA02
             |              |--P. pickeringi Piper 2006BA12
             |              |--P. rephaim Ramsay 1885F71
             |              `--P. selestiae Mackness 1995LA02
             `--Vombatidae [Phascolomidae, Vombatoidea]BA12
                  |  i. s.: Wombatula Troughton 1941R64
                  |           `--*W. gillespiei (De Vis 1900) [=Phascolomys gillespiei]R64
                  |         ‘Sarcophilus’ prior De Vis 1883WHR82
                  |--Rhizophascolonus Stirton, Tedford & Woodburne 1967LA02
                  |    `--*R. crowcrofti Stirton, Tedford & Woodburne 1967LA02
                  `--+--Warendja Hope & Wilkinson 1982LA02
                     |    |--*W. wakefieldi Hope & Wilkinson 1982LA02
                     |    `--W. encorensis Brewer et al. 2007BA12
                     `--+--Vombatus Geoffroy 1803LA02
                        |    |--*V. ursinus (Shaw 1800)LA02
                        |    |--V. hacketti (Glauert 1910)LA02
                        |    `--V. hirsutus (see below for synonymy)W78
                        `--+--Ramsayia Tate 1951LA02
                           |    |--*R. magna (Owen 1872)LA02
                           |    `--R. lemleyi (Archer 1976)LA02
                           `--+--Lasiorhinus Gray 1863LA02
                              |    |--*L. latifrons (Owen 1845)LA02
                              |    |    |--L. l. latifronsR64
                              |    |    `--L. l. barnardi Longman 1939R64
                              |    |--L. angustidens (De Vis 1891)LA02
                              |    |--L. gillespieiUSDI77
                              |    `--L. krefftiiCB-E04
                              `--+--Phascolonus Owen 1872LA02
                                 |    `--*P. gigas (Owen 1858)LA02 [=Phascolomys (Phascolonus) gigasF71]
                                 `--PhascolomysLA02
                                      |--P. curvirostris Owen 1885F71
                                      |--P. medius Owen 1872LA02
                                      |--P. mitchelliB97
                                      |--P. ursinusA14
                                      `--‘Phascolomis’ vombatus Leach 1815R64

Thylacoleo Gervais 1852LA02 [incl. Plectodon Krefft 1870 non Carpenter 1864AD82, Schizodon Stutchbury 1853 nec Agassiz 1829 nec Waterhouse 1842AD82, Thylacopardus Owen 1888AD82; Thylacoleoninae]

*Thylacoleo carnifex Owen 1858LA02, AD82, LA02 [incl. Mylodon australis Krefft 1870AD82, Thylacopardus australis Owen 1888AD82, Prochaerus celer De Vis 1886AD82, Thylacoleo oweni McCoy 1876AD82, Thylacoleo robustus Krefft 1872AD82]

Vombatus hirsutus [incl. V. mitchellii, V. parvus, Phascolomys platyrhinus, P. pliocenus McCoy 1866, Vombatus pliocenus, V. thompsoni]W78

*Type species of generic name indicated

References

[A14] Anderson, W. 1914. Notes on the occurrence of the sand-rock containing bones of extinct species of marsupials (emu, kangaroo, wombat, etc.) on King Island, Bass Strait, Tasmania. Records of the Australian Museum 10 (9): 275–283.

[AD82] Archer, M., & L. Dawson. 1982. Revision of marsupial lions of the genus Thylacoleo Gervais (Thylacoleonidae, Marsupialia) and thylacoleonid evolution in the Late Cainozoic. In: Archer, M. (ed.) Carnivorous Marsupials vol. 2 pp. 477–494. Royal Zoological Society of New South Wales.

[B07] Black, K. 2007. Maradidae: a new family of vombatimorphian marsupial from the late Oligocene of Riversleigh, northwestern Queensland. Alcheringa 31 (1): 17–32.

[BA12] Black, K. H., M. Archer, S. J. Hand & H. Godthelp. 2012. The rise of Australian marsupials: a synopsis of biostratigraphic, phylogenetic, palaeoecologic and palaeobiogeographic understanding. In: Talent, J. A. (ed.) Earth and Life: Global biodiversity, extinction intervals and biogeographic perturbations through time pp. 983–1078. Springer.

[B97] Broom, R. 1897. On the comparative anatomy of the organ of Jacobson in marsupials. Proceedings of the Linnean Society of New South Wales 21 (4): 591–623, pls 41–48.

[CB-E04] Cardillo, M., O. R. P. Bininda-Emonds, E. Boakes & A. Purvis. 2004. A species-level phylogenetic supertree of marsupials. Journal of Zoology 264: 11–31.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.

[LA02] Long, J., M. Archer, T. Flannery & S. Hand. 2002. Prehistoric Mammals of Australia and New Guinea: One Hundred Million Years of Evolution. University of New South Wales Press: Sydney.

[R64] Ride, W. D. L. 1964. A list of mammals described from Australia between the years 1933 and 1963 (comprising newly proposed names and additions to the Australian faunal list). Australian Mammal Society Bulletin 7 (Suppl.): 1–15.

[USDI77] United States Department of the Interior. 1977. Endangered and threatened wildlife and plants—republication of list of species. Federal Register 42: 36420–36431.

[WHR82] Wells, R. T., D. R. Horton & P. Rogers. 1982. Thylacoleo carnifex Owen (Thylacoleonidae): marsupial carnivore?. In: Archer, M. (ed.) Carnivorous Marsupials vol. 2 pp. 573–585. Royal Zoological Society of New South Wales.

[W78] Wilkinson, H. E. 1978. Synonymy of the fossil wombat Vombatus pliocenus (McCoy) with the living species Vombatus hirsutus (Perry). Memoirs of National Museum Victoria 39: 93–100.

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