European shot borer Xyleborus dispar in plum tree, from here.

Belongs within: Hylesininae.

Xyleborus is a cosmopolitan genus of xylomycetophagous (i.e. feeding on fungi growing on wood) bark beetles. Members of this genus are consanguineously polygynous, with haploid flightless males (Wood 1986).

<==Xyleborini [Webbinae, Xylebori, Xyleborina]
    |--Xyleborites Wickham 1913W86
    |    `--*X. longipennis Wickham 1913W86
    |--Sampsonius Eggers 1935W86
    |    `--*S. sexdentatusW86
    |--Schedlia Browne 1950W86
    |    `--*S. sumatrana [=Xyleborus sumatranus]W86
    |--Dryocoetoides Hopkins 1915W86
    |    `--D. capucinus (Eichh. 1868)W86, FS90 [=Xyleborus capucinusW86; incl. *D. guatemalensisW86]
    |--Leptoxyleborus Wood 1980W86
    |    `--*L. sordicauda [=Phloeotrogus sordicauda]W86
    |--Theoborus Hopkins 1915W86
    |    `--*T. theobromaeW86
    |--Taurodemus Wood 1980W86
    |    `--*T. sharpi [=Xyleborus sharpi]W86
    |--Mesoscolytus Broun 1904W86
    |    `--*M. inurbanusW86
    |--Hadrodemius Wood 1980W86
    |    `--*H. globus [=Xyleborus globus]W86
    |--Taphrodasus Wood 1980W86
    |    `--*T. percorthylus [=Xyleborus percorthylus]W86
    |--Cryptoxyleborus Schedl 1937W86
    |    `--*C. naevusW86
    |--Anisandrus Ferrari 1867GC-P14, W86
    |    `--*A. disparW86, GC-P14 [=Xyleborus (Apate) disparW86]
    |--Euwallacea Hopkins 1915W86
    |    |--*E. wallacei [=Xyleborus wallacei]W86
    |    `--E. validus (Eichhoff 1875)A02
    |--Cnestus Sampson 1911 [incl. Tosaxyleborus Murayama 1950]W86
    |    |--*C. magnusW86
    |    `--C. murayamai [incl. *Tosaxyleborus pallidipennis]W86
    |--Coptoborus Hopkins 1915 [incl. Streptocranus Schedl 1939]W86
    |    |--C. vespatorius [=Xyleborus vespatorius; incl. *C. emarginatus]W86
    |    `--C. mirabilis [=*Streptocranus mirabilis]W86
    |--Arixyleborus Hopkins 1915 [incl. Xyleboricus Eggers 1923]W86
    |    |--*A. rugosipesW86
    |    `--A. canaliculatus [=*Xyleboricus canaliculatus]W86
    |--Terminalinus Hopkins 1915 [incl. Kelantanius Nunberg 1961]W86
    |    |--*T. terminaliaeW86
    |    `--T. punctatopilosus [=Xyleborus punctatopilosus, *Kelantanius punctatopilosus]W86
    |--Eccoptopterus Motschulsky 1863 (see below for synonymy)W86
    |    |--E. spinosus [=Scolytus spinosus; incl. *Eccoptopterus sexspinosus]W86
    |    `--E. gracilipes [=*Platydactylus gracilipes, *Eurydactylus gracilipes]W86
    |--Xyleborinus Reitter 1913M94
    |    |--*X. saxeseni (Ratzeburn 1837) [=Bostrichus saxeseni]M94
    |    `--X. alniA02
    |--Ambrosiodmus Hopkins 1915 [incl. Browneia Nunberg 1963, Phloeotrogus Motschulsky 1863 (nom. rej.)]W86
    |    |--*A. tachygraphus [=Xyleborus tachygraphus]W86
    |    |--A. obliquecaudata [=*Phloeotrogus oliquecaudata]W86
    |    `--A. obliquus [=Pityophthorus obliquus; incl. Xyleborus illepidus, *Browneia illepida]W86
    |--Amasa Lea 1894 [incl. Anaxyleborus Wood 1980, Pseudoxyleborus Eggers 1930]W86
    |    |--*A. thoracicusW86
    |    |--*Pseudoxyleborus’ beesoniW86
    |    `--A. truncatus (Erichs. 1842)W86, M87 [=Tomicus truncatusW86, *Anaxyleborus truncatusW86]
    |--Coptodryas Hopkins 1915 [incl. Adryocoetes Eggers in Schedl 1952 (n. n.), Microperus Wood 1980]W86
    |    |--*C. confusaW86
    |    |--C. pullus [=Xyleborus pullus; incl. *Adryocoetes nitidus (n. n.)]W86
    |    `--C. theae [=Xyleborus theae, *Microperus theae]W86
    |--Webbia Hopkins 1915 [incl. Prowebbia Browne 1962, Pseudowebbia Browne 1961, Xelyborus Schedl 1939 (n. n.)]W86
    |    |--*W. dipterocarpiW86
    |    |--W. paboW86
    |    |--W. subuculae [=*Prowebbia subuculae]W86
    |    `--W. trepanicauda [=Xyleborus trepanicauda, *Pseudowebbia trepanicauda]W86
    |--Xylosandrus Reitter 1913 [incl. Apoxyleborus Wood 1980]W86
    |    |--*X. morigerus [=Xyleborus morigerus]W86
    |    |--X. compactus (Eichhoff 1875)A02
    |    |--X. mancus [=Xyleborus mancus, *Apoxyleborus mancus]W86
    |    `--X. retususW86
    `--Xyleborus Eichhoff 1864W86 (see below for synonymy)
         |--*X. monographus [=Bostrichus monographus]W86
         |--X. affinis Eichh. 1867 [incl. X. retusicollis Zimm. 1868]FS90
         |--X. bidentatus [=Phloeotrogus bidentatus; incl. *Boroxylon stephegynis]W86
         |--X. confusus Eichh. 1867FS90
         |--X. cryptophagus [=Bostrichus cryptographus, *Heteroborips cryptographus]W86
         |--X. ferrugineusRD77
         |--X. fuscatus Eichh. 1867FS90
         |--X. kalshoveni [=*Notoxyleborus kalshoveni]W86
         |--X. meuseli [=*Xyleborips meuseli]W86
         |--X. morstattaBW64
         |--X. posticus Eichh. 1868FS90
         |--X. prelaetus [incl. *Cyclorhipidion pelliculosum]W86
         |--X. pubescensW86
         |--X. saxeseniV09
         |--X. solidus Eichh. 1868M87
         |--X. spathipennisW86
         |--X. subcostatus [incl. *Progenius fleutiauxi]W86
         `--X. volvulus [=Bostrichus volvulus; incl. X. guanajuatensis, *Anaeretus guanajuatensis]W86

Eccoptopterus Motschulsky 1863 [incl. Eurydactylus Hagedorn 1909 non LaFerté Sénectère 1851, Platydactylus Eichhoff 1886]W86

Xyleborus Eichhoff 1864W86 [incl. Anaertus Dugès 1887A02, Boroxylon Hopkins 1915W86, Cyclorhipidion Hagedorn 1912W86, Heteroborips Reitter 1913W86, Notoxyleborus Schedl 1934W86, Progenius Blandford 1896W86, Xyleborips Reitter 1913W86]

*Type species of generic name indicated


[A02] Anderson, R. S. 2002. Curculionidae Latreille 1802. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 722–815. CRC Press.

[BW64] Booth, C., & J. M. Waterston. 1964. Calonectria rigidiuscula. C.M.I. Descriptions of Pathogenic Fungi and Bacteria 21.

[FS90] Fleutiaux, E., & A. Sallé. 1890. Liste des coléoptères de la Guadeloupe et descriptions d’espèces nouvelles. Annales de la Société Entomologique de France, 6e série 9: 351–484.

[GC-P14] Gillett, C. P. D. T., A. Crampton-Platt, M. J. T. N. Timmermans, B. H. Jordal, B. C. Emerson & A. P. Vogler. 2014. Bulk de novo mitogenome assembly from pooled total DNA elucidates the phylogeny of weevils (Coleoptera: Curculionoidea). Molecular Biology and Evolution 31 (8): 2223–2237.

[M87] Masters, G. 1887. Catalogue of the described Coleoptera of Australia. Part VI. Proceedings of the Linnean Society of New South Wales, series 2, 1 (4): 979–1036.

[M94] May, B. M. 1994. An introduction to the immature stages of Australian Curculionoidea. In: Zimmerman, E. C. Australian Weevils (Coleoptera: Curculionoidea) vol. 2. Brentidae, Eurhynchidae, Apionidae and a chapter on immature stages by Brenda May pp. 365–728. CSIRO Australia.

[RD77] Richards, O. W., & R. G. Davies. 1977. Imms’ General Textbook of Entomology 10th ed. vol. 2. Classification and Biology. Chapman and Hall: London.

[V09] Verdcourt, B. (ed.) 2009. Additions to the wild fauna and flora of the Royal Botanic Gardens, Kew. XXVI. Miscellaneous records. Kew Bulletin 64 (1): 183–194.

[W86] Wood, S. L. 1986. A reclassification of the genera of Scolytidae (Coleoptera). Great Basin Naturalist Memoirs 10: 1–126.

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