Common sun beetle Amara aenea, copyright Entomart.

Belongs within: Harpalinae.

The Amarini are a group of ground beetles with a head distinctly wider than long, and short, stout mandibles.

<==Zabrini [Amarini]
    |--Pseudamara Lindroth 1968 [incl. Disamara Lindroth 1976, Harpaloamara Hieke 1995]BB01
    |    `--P. arenaria LeConte 1848BB01 [=Disamara arenariaB90]
    |--Zabrus Clairville 1806BB01
    |    |--Z. gibbus [=Harpalus (Zabrus) gibbus]G20
    |    `--Z. triniiC01
    `--Amara Bonelli 1810BB01
         |  i. s.: A. aeneaC01
         |         A. aurichalceaC01
         |         A. convexaHH96
         |         A. equestrisC01
         |         A. fodinaeC01
         |         A. hyperboreaBB01
         |         A. municipalisC01
         |         A. praetermissaC01
         |         A. similataC01
         |         A. solskyiC01
         |--A. (Amara) [incl. Celia Zimmermann 1832, Isopleurus Kirby 1837]BB01
         |    |--A. (A.) aurataBB01
         |    |--A. (A.) californica [=A. (Celia) californica]BB01
         |    |--A. (A.) fulva [=Harpalus (A.) fulvus]G20
         |    |--A. (A.) lunicollisBB01
         |    `--A. (A.) quenseliBB01
         |--A. insignis groupBB01
         |    |--A. insignis Dejean 1831 [=A. (Bradytus) insignis]BB01
         |    `--A. insularis Horn 1875 [=A. (Bradytus) insularis]BB01
         |--A. (Bradytus Stephens 1827) [incl. Leiocnemis Zimmermann 1832]BB01
         |    |--A. (B.) apricariaBB01
         |    `--A. (B.) colvillensis Lindroth 1968BB01
         |--A. (Curtonotus Stephens 1828) [=Cyrtonotus]BB01
         |    `--A. (C.) aulicaBB01
         |--A. (Neopercosia Hieke 1978)BB01
         |    `--A. (N.) fortis LeConte 1880BB01
         |--A. (Percosia Zimmermann 1832)BB01
         |    `--A. (P.) obesa Say 1823BB01
         |--A. (Reductocelia Lafer 1989)BB01
         `--A. (Zezea Csiki 1929) [incl. Triaena LeConte 1848 non Hübner 1818]BB01
              `--A. (Z.) pallipesBB01

*Type species of generic name indicated


[BB01] Ball, G. E., & Y. Bousquet. 2001. Carabidae Latreille, 1810. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 32–132. CRC Press: Boca Raton.

[B90] Bell, R. T. 1990. Insecta: Coleoptera Carabidae adults and larvae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1053–1092. John Wiley & Sones: New York.

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[HH96] Husband, R. W., & D. O. Husband. 1996. A new species of Eutarsopolipus (Acari: Podapolipidae) from Amara californica Dejean (Coleoptera: Carabidae) from California. Proceedings of the Entomological Society of Washington 98 (3): 465–470.

Leave a comment

Your email address will not be published. Required fields are marked *