Zoila friendii, copyright S. & A. Nappo.

Belongs within: Cypraeidae.

Zoila is a genus of medium- to large-sized cowries currently restricted to coasts of Australia, though fossils indicate a wider distribution in the western Pacific in the past. Members of this genus are direct developers, without a planktonic larval stage (Meyer 2003).

<==Zoila Jousseaume 1884 [Zoilinae]BR05
    |  i. s.: *Z. scottii (Broderip 1831) [=Cypraea scottii]BR17
    |         Z. alabaster Mont & Lorenz 2013WBS-S15
    |         Z. gigas (McCoy 1867) [=Cypraea (Zoila) gigas]WG71
    |         Z. mariellae (Raybaudi 1983) [=Cypraea mariellae]W93
    |         Z. perlae (Lopez & Chiang 1975) [=Cypraea perlae]W93
    |--Z. marginata Gaskoin 1849M03 [=Cypraea (Zoila) marginataWG71]
    |    |--Z. m. marginata [incl. Cypraea marginata raybaudii Lorenz 1992]W93
    |    |--Z. m. albanyensis (Raybaudi 1985) [=Cypraea marginata albanyensis]W93
    |    |--Z. m. consueta (Biraghi 1993) [=Cypraea marginata consueta]W93
    |    `--Z. m. ketyana Raybaudi 1978 [=Cypraea marginata ketyana; incl. C. intermedia Bodoni 1986 (n. n.)]W93
    |--Z. rosselli Cotton 1948M03 (see below for synonymy)
    |--Z. venusta Sowerby 1846M03 (see below for synonymy)
    |    |--Z. v. venustaBC01
    |    `--Z. v. episema (Iredale 1939)BC01 (see below for synonymy)
    `--+--Z. decipiens Smith 1880M03 [=Cypraea (Zoila) decipiensWG71]
       |--Z. eludens Raybaudi 1991M03 [=Cypraea eludensW93]
       `--+--Z. friendii Gray 1831M03 [=Cypraea (Zoila) friendiiWG71]
          |    |--Z. f. friendii (see below for synonymy)W93
          |    |--Z. f. jeaniana Cate 1968MG-H11 [=Cypraea friendii jeanianaW93; incl. C. aurata Raybaudi 1978W93]
          |    |--Z. ‘jenaiana’ sherylae Raybaudi 1990M03
          |    `--Z. f. vercoi Schilder 1930W93 (see below for synonymy)
          `--Z. thersites Gaskoin 1849 [=Cypraea (Zoila) friendii thersites]M03

Zoila friendii friendii Gray 1831 [incl. C. friendii f. dorata Raybaudi 1992, C. friendii f. maxima Raybaudi 1992, C. friendii f. nana Raybaudi 1992]W93

Zoila friendii vercoi Schilder 1930W93 [=Cypraea friendii vercoiW93; incl. C. thersites contraria Iredale 1935W93, C. friendii f. contraria WG71]

Zoila rosselli Cotton 1948M03 [=Cypraea (Zoila) rosselliWG71; incl. Z. rosselli albosignata Raybaudi 1985 (n. n.)W93, Z. rosselli edingeri Raybaudi 1990W93]

Zoila venusta Sowerby 1846M03 [=Cypraea (Zoila) venustaWG71; incl. C. bakeri Gatliff 1916W93, C. catei Schilder 1963WG71, Zoila venusta orientalis Raybaudi 1992W93, C. roseopunctata Melvill 1888W93, C. thatcheri Cox 1869W93]

Zoila venusta episema (Iredale 1939)BC01 [=Cypraea venusta episemaWG71; incl. Z. venusta sorrentensis Schilder 1963BC01, C. venusta sorrentensisWG71]

*Type species of generic name indicated


[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1-2): 1-397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[M03] Meyer, C. P. 2003. Molecular systematics of cowries (Gastropoda: Cypraeidae) and diversification patterns in the tropics. Biological Journal of the Linnean Society 79: 401–459.

[WBS-S15] Willan, R. C., C. Bryce & S. M. Slack-Smith. 2015. Kimberley marine biota. Historical data: molluscs. Records of the Western Australian Museum Supplement 84: 287–343.

[W93] Wilson, B. 1993. Australian Marine Shells vol. 1. Prosobranch Gastropods. Part One. Odyssey Publishing: Kallaroo (Western Australia).

[WG71] Wilson, B. R., & K. Gillett. 1971. Australian Shells: illustrating and describing 600 species of marine gastropods found in Australian waters. A. H. & A. W. Reed: Sydney.

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